Glossary of botanical terms used in the Poaceae
Adapted from the glossary in Flora of Ethiopia and Eritrea, vol. 7 (1995).
aristate - with an awn
aristulate - diminutive of aristate
auricle - an earlike lobe or appendage at the junction of leaf sheath and blade
auriculate - with an auricle
awn - a bristle arising from a spikelet part
callus - a hard projection at the base of a floret, spikelet, or inflorescence segment, indicating a disarticulation point
caryopsis - a specialized dry fruit characteristic of grasses, in which the seed and ovary wall have become united
collar - pale or purplish zone at the junction of leaf sheath and blade
column - the lower twisted portion of a geniculate awn, or the part below the awn branching-point in Aristideae
compound - referring to inflorescences made up of a number of small constituent inflorescences (as in some Andropogoneae), or a raceme with some secondary branching
culm - the flowering stem of a grass plant
culm sheath (bamboos) - modified, often non-photosynthetic, culm leaf with an expanded sheath and much reduced blade, usually deciduous as the culm matures
diffuse (bamboos) - culms arising singly from long slender rhizomes.
extravaginal - branching in which the young shoot breaks through the base of the leaf sheath
floret - the individual unit of a spikelet, comprising a lemma and palea with enclosed reproductive organs
glume - one of a pair of empty scales at the base of a grass spikelet
heterogamous spikelets - the paired spikelets found in most Andropogoneae, where one spikelet of the pair is sessile and produces a caryopsis, and the other spikelet is pedicelled, of different form, and staminate or sterile
hilum - the scar on the caryopsis marking the site of the attachment of the pericarp and testa, found on the opposite side from the embryo
homogamous spikelets - in Andropogoneae the paired spikelets sometimes present at the base of the raceme, of similar appearance and not producing any caryopses, often resembling the pedicelled spikelets or assuming a protective involucral function
intravaginal - branching in which the young shoot grows up inside the leaf sheath, emerging at the sheath mouth
iterauctant (bamboos) - inflorescence with pseudospikelets with glumes subtending axillary buds capable of partial or extensive spikelet ramification
leaf blade - the distal expanded part of a grass leaf
leaf sheath - the basal part of the grass leaf which normally encloses a culm internode
lemma - the lower of the two bracts enclosing the grass flower and together with the palea comprising a floret
leptomorph (bamboos) - rhizome monopodial, elongated, more slender than culms
ligule - a membrane or line of hairs on the inner (adaxial) side of the junction of the leaf sheath and leaf blade; bamboos sometimes have an external ligule on the abaxial side of the junction
lodicule - a small scale-like or fleshy structure at the base of the stamens in a grass floret, usually 2 in each floret (often 3 or more in bamboos); they swell at anthesis, causing the floret to gape open
oral setae - marginal setae inserted at junction of leaf sheath and blade, on the auricles when these are present
pachymorph (bamboos) - rhizome sympodial, thicker than culms
palea - the upper and inner scale of the grass floret which encloses the grass flower, usually 2-keeled
panicle - in grasses, an inflorescence in which the primary axis bears branched secondary axes with pedicellate spikelets
pedicel - in grasses, the stalk of a single spikelet within an inflorescence
peduncle - the stalk of a raceme or cluster of spikelets
pluricaespitose (bamboos) - culms arising in a series of clusters along a long slender rhizome
prophyll - in grasses, a 2-keeled, hyaline, modified leaf, placed within a leaf sheath on the adaxial side of a branch
pseudopetiole - the narrow basal portion of some leaf blades, resembling a petiole
pseudospikelet (bamboos) - spikelet in which the outer glumes or bracts subtend axillary buds which can develop to form lateral spikelets or branches
raceme - in grasses, an unbranched axis bearing spikelets; racemes may be solitary, digitate, or scattered
raceme base - short stalk beneath the individual racemes of a pair in some Andropogoneae
raceme pair - pairs of racemes supported by spatheoles in the compound panicles of some Andropogoneae
rachilla - the central axis of the spikelet which bears the florets
rachilla extension - a prolongation of the rachilla beyond the uppermost (or single) floret
rachis - the axis of a raceme
secondary spathe - spathe supporting a second tier of branching within the compound panicle of some Andropogoneae
semelauctant (bamboos) - inflorescence with glumes not subtending viable buds or branches
sinus - the space between two projecting lobes or teeth
spathate - with spathes
spathe - a bract or modified bladeless leaf subtending the inflorescence or part of it
spatheole - the uppermost spathe supporting the racemes within the compound inflorescence of some Andropogoneae
spikelet - the basic unit of a grass inflorescence; usually composed of two glumes and one or more florets on a rachilla
spikelet pair - the arrangement of one sessile and one pedicelled spikelet arising from the same node characteristic of the Andropogoneae
tiller - a leafy non-flowering shoot
triad - a group of three spikelets borne together
unicaespitose (bamboos) - culms all arising in a single clump from pachymorph rhizomes
Figure 1. 1-4. Parts of a typical grass (based on Bromus). —1. Portion of leaf. —2. Spikelet. —3. Floret. —4. Flower. 5-7. Compound panicle and raceme pair structure in Andropogoneae (based on Hyparrhenia). —5. Compound panicle with primary and secondary tiers. —6. Compound panicle with primary tier showing raceme pair. —7. Diagram of raceme pair structure. Drawn by Yevonn Wilson-Ramsey.