Brassicaceae Genera of the World

Ihsan Al-Shehbaz


Acanthocardanum

Perennial; shrubs or subshrubs; terrestrial; thorns and spines present; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; oblong to elliptic; margin and apex entire. Free, or united. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovate or obovate; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed); fruit appendages wings. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading.

Iran.


Aethionema

Acanthocardanum Thell.; Eunomia DC.; Iberidella Boiss.; Moriera Boiss.

Annual or biennial, or perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines present, or absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow, or pink (or lavender), or purple (or violet); obovate, or oblong to elliptic; margin and apex entire. Sepals ascending, or erect but not connivent, or spreading; free. Stamens six; lower part of filaments and petal claws without anything, or filaments with a wing, tooth, or appendage; filaments of median stamens free, or united at least along lower third; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); oblong, or elliptic, or ovate or obovate, or orbicular; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds), or indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages wings. Ovules/seeds per ovary/fruit two, or three to five, or six to ten; Seeds uniseriate (in one row), or aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate, or opposite. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Afghanistan, or Algeria, or Armenia, or Azerbaijan, or Egypt, or Georgia, or Iran, or Iraq, or Israel, or Jordan, or Kazakhstan, or Lebanon, or Morocco, or Pakistan, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or Uzbekistan, or Greece, or Italy, or Bulgaria, or Albania, or Crete, or Czech R, or France, or Germany, or Switzerland, or Spain, or Hungary, or Yugoslavia, or Romania, or Sardinia, or Sicily, or Austria, or Croatia, or Slovakia, or Slovenia, or Kyrgyzstan, or Czech R, or Slovakia, or Slovenia, or Algeria, or Bosnia & Herz, or Macedonia.


Alliaria

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or Armenia, or Azerbaijan, or China, or Cyprus, or Georgia, or India, or Iran, or Iraq, or Kazakhstan, or Kyrgyzstan, or Lebanon, or Nepal, or Pakistan, or Russia, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or Uzbekistan, or Algeria, or Morocco, or Tunisia, or Estonia, or Latvia, or Lithuania, or Belarus, or Moldova, or Ukraine, or Czech R, or Slovakia, or Slovenia, or Canada, or United States, or Tunisia, or Algeria, or New Zealand, or Ireland, or United Kingdom, or Greece, or Crete, or Portugal, or Spain, or Albania, or Austria, or Belgium, or Bosnia & Herz, or Bulgaria, or Corsica, or Croatia, or Denmark, or Finland, or France, or Germany, or Netherlands, or Hungary, or Italy, or Macedonia, or Norway, or Poland, or Romania, or Sardinia, or Sicily, or Sweden, or Switzerland, or Yugoslavia.


Alyssoides

Lutzia Gandoger; Vesicaria Adans.

Perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; oblong to elliptic; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); orbicular; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds), or indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten; Seeds biseriate (in two rows); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Armenia, or Georgia, or Turkey, or Crete, or Azerbaijan, or Greece, or Albania, or Bulgaria, or France, or Italy, or Yugoslavia, or Romania, or Bosnia & Herz, or Croatia, or Macedonia.


Alyssopsis

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; orbicular; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Azerbaijan, or Iran, or Turkmenistan.


Alyssum

Gamosepalum Hausskn.; Meniocus Desv.; Odontarrhena C.A. Mey.; Psilonema C.A. Mey.; Ptilotrichon C.A. Mey.; Takhtajaniella V.E. Avet.; Triplopetalum Nyarady.

Annual or biennial, or perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines present, or absent; hairs (trichomes) a mixture of simple and branched, or stellate (star-shaped) and sessile; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow, or pink (or lavender); obovate, or oblong to elliptic; margin and apex entire. Sepals ascending, or erect but not connivent, or erect and connivent; free, or united. Stamens six; lower part of filaments and petal claws without anything, or filaments with a wing, tooth, or appendage; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic, or orbicular, or globose (spherical); latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none, or wings. Ovules/seeds per ovary/fruit one, or two, or three to five, or six to ten; Seeds biseriate (in two rows), or aseriate (not in rows: when fruit has one or a few seeds); wing present, or absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or Armenia, or Azerbaijan, or China, or Cyprus, or Georgia, or India, or Iran, or Iraq, or Israel, or Jordan, or Kazakhstan, or Kuwait, or Kyrgyzstan, or Lebanon, or Mongolia, or Pakistan, or Russia, or Saudi Arabia, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or Uzbekistan, or Yemen, or Argentina, or Chile, or Estonia, or Latvia, or Lithuania, or Belarus, or Moldova, or Egypt, or Ukraine, or Czech R, or Slovakia, or Slovenia, or Canada, or United States, or Australia, or Algeria, or Libya, or Morocco, or Tunisia, or New Zealand, or Canary Islands, or South Africa, or Ireland, or United Kingdom, or Crete, or Greece, or Portugal, or Spain, or Albania, or Austria, or Belgium, or Bosnia & Herz, or Bulgaria, or Corsica, or Croatia, or Denmark, or Finland, or France, or Germany, or Netherlands, or Hungary, or Italy, or Macedonia, or Norway, or Poland, or Romania, or Sardinia, or Sicily, or Sweden, or Switzerland, or Yugoslavia, or Balearic Islds.


Ammosperma

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or purple (or violet); obovate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit more than 100; Seeds biseriate (in two rows); wing present; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Algeria, or Libya, or Tunisia.


Anastatica

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate, or oblong to elliptic; margin and apex entire. Sepals erect but not connivent, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); globose (spherical), or ovoid or obovoid; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick; indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages wings. Ovules/seeds per ovary/fruit three to five; Seeds uniseriate (in one row); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Bahrain, or Iran, or Iraq, or Israel, or Jordan, or Kuwait, or Oman, or Pakistan, or Qatar, or Saudi Arabia, or Turkey, or Yemen, or Egypt, or Libya, or Algeria, or Morocco, or Sudan, or Eritrea, or Mauritania, or Mali, or Ethiopia.


Anchonium

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple, or at least some dendritic (tree-like); multicellular glands present; not a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are bracteate throughout or at least in lower half. Petals distinctly longer than sepals; yellow, or purple (or violet); obovate; margin and apex entire. Sepals erect but not connivent, or erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens united at least along lower third; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); lanceolate or oblanceolate, or ellipsoid; terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall woody or corky and thick; breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten; Seeds uniseriate (in one row); wing present, or absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Armenia, or Azerbaijan, or Georgia, or Iran, or Iraq, or Lebanon, or Syria, or Turkey.


Andrzeiowskia

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or oblong; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages wings, or horns. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Greece, or Lebanon, or Turkey.


Anelsonia

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves absent; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); obovate, or oblong to elliptic; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); lanceolate or oblanceolate, or elliptic; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds biseriate (in two rows); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

United States.


Aphragmus

Lignariella Baehni; Staintoniella H. Hara.

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple, or a mixture of simple and branched; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or palmately lobed. Inflorescences racemes that are bracteate throughout or at least in lower half. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet); obovate, or orbicular; margin and apex entire. Sepals ascending, or erect but not connivent, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or lanceolate or oblanceolate, or oblong, or elliptic, or ovate or obovate, or ellipsoid; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore at least 1.5 mm long, usually much longer, or absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit two, or three to five, or six to ten, or eleven to 20; Seeds uniseriate (in one row), or biseriate (in two rows), or aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Afghanistan, or Bhutan, or Canada, or China, or India, or Mongolia, or Nepal, or Pakistan, or Russia, or Tajikistan, or United States, or Kyrgyzstan.


Aplanodes

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or purple (or violet); obovate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything, or filaments with a wing, tooth, or appendage; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

South Africa, or Lesotho.


Arabidella

Lemphoria O.E. Schulz; Micromystria O.E. Schulz; Pseudoarabidella O.E. Schulz.

Annual or biennial, or perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate, or solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals, or subequalling sepals; white, or yellow, or pink (or lavender); obovate, or orbicular; margin and apex entire. Sepals spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; angustiseptate (flattened at a right angle to septum), or latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50, or 51 to 100; Seeds uniseriate (in one row), or biseriate (in two rows); wing present, or absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Australia.


Arabidopsis

Cardaminopsis Hayek; Hylandra A. Löve; Stenophragma Celak.

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals; white, or pink (or lavender), or purple (or violet); obovate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50, or 51 to 100; Seeds uniseriate (in one row); wing present, or absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or Armenia, or Azerbaijan, or China, or Cyprus, or Georgia, or India, or Iran, or Kazakhstan, or Kyrgyzstan, or Lebanon, or Morocco, or Pakistan, or Russia, or Saudi Arabia, or Syria, or Tajikistan, or Turkey, or Uzbekistan, or Estonia, or Latvia, or Lithuania, or Belarus, or Moldova, or Ukraine, or Czech R, or Slovakia, or Slovenia, or United States, or Australia, or Tasmania, or Tunisia, or Libya, or Morocco, or Algeria, or Tanzania, or Kenya, or Uganda, or Ethiopia, or Congo Democratic R, or Canary Islands, or Madeira, or Somalia, or Sudan, or New Zealand, or Cape Verde, or Taiwan, or South Africa, or Argentina, or Uruguay, or Ireland, or United Kingdom, or Djibouti, or Rwanda, or Crete, or Greece, or Portugal, or Spain, or Albania, or Austria, or Belgium, or Bosnia & Herz, or Bulgaria, or Corsica, or Croatia, or Denmark, or Balearic Islds, or Finland, or France, or Germany, or Netherlands, or Hungary, or Iceland, or Italy, or Macedonia, or Norway, or Poland, or Romania, or Sardinia, or Sicily, or Sweden, or Switzerland, or Yugoslavia.


Arabis

Parryodes Jafri.

Annual or biennial, or perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively malpighiaceous (2-branched, sessile, medifixed), or stellate and stalked; multicellular glands absent; not a weed, or a weed. Stem leaves present; base sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are bracteate throughout or at least in lower half, or racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet); obovate, or oblong to elliptic, or orbicular; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50, or 51 to 100; Seeds uniseriate (in one row), or biseriate (in two rows); wing present, or absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or Armenia, or Azerbaijan, or Bhutan, or China, or Cyprus, or India, or Iran, or Iraq, or Israel, or Japan, or Jordan, or Kazakhstan, or Korea, or Kyrgyzstan, or Lebanon, or Mongolia, or Nepal, or Pakistan, or Russia, or Saudi Arabia, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or Uzbekistan, or Yemen, or Estonia, or Latvia, or Lithuania, or Belarus, or Moldova, or Egypt, or Ukraine, or Czech R, or Slovakia, or Slovenia, or Canada, or United States, or Greenland, or Algeria, or Libya, or Morocco, or Tunisia, or Kenya, or Tanzania, or Uganda, or Rwanda, or Congo Democratic R, or Eritrea, or Ethiopia, or Madeira, or Canary Islands, or Taiwan, or Ireland, or United Kingdom, or Rwanda, or Crete, or Greece, or Portugal, or Spain, or Albania, or Austria, or Belgium, or Bosnia & Herz, or Bulgaria, or Corsica, or Croatia, or Denmark, or Finland, or France, or Germany, or Netherlands, or Hungary, or Italy, or Macedonia, or Norway, or Poland, or Romania, or Sardinia, or Sicily, or Sweden, or Switzerland, or Yugoslavia.


Arcyosperma

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); obovate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Bhutan, or India, or Pakistan.


Armoracia

Perennial; herbs; terrestrial, or aquatic; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate, or oblong to elliptic; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); oblong, or orbicular; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds biseriate (in two rows); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Armenia, or Azerbaijan, or Georgia, or Japan, or Kazakhstan, or Russia, or Tajikistan, or Turkey, or Uzbekistan, or Estonia, or Latvia, or Lithuania, or Belarus, or Kyrgyzstan, or Moldova, or Ukraine, or Czech R, or Slovakia, or Slovenia, or Mexico, or Canada, or United States, or Libya, or New Zealand, or Ireland, or United Kingdom, or Spain, or Albania, or Austria, or Belgium, or Bosnia & Herz, or Bulgaria, or Denmark, or Finland, or France, or Germany, or Switzerland, or Netherlands, or Hungary, or Italy, or Yugoslavia, or Norway, or Poland, or Romania.


Aschersoniodoxa

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves absent; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or lanceolate or oblanceolate, or oblong; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina, or Bolivia, or Peru.


Asperuginoides

Buchingera Boiss. & Hohen.

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) stellate (star-shaped) and sessile; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are bracteate throughout or at least in lower half. Petals subequalling sepals; white; obovate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); oblong, or orbicular; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages glochidiate trichomes. Ovules/seeds per ovary/fruit one, or two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit reflexed (pendulous). Fruit segmentation not segmented.

Armenia, or Iran, or Kazakhstan, or Pakistan, or Tajikistan, or Turkmenistan, or Uzbekistan.


Asta

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate, or oblong to elliptic; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); oblong, or ovate or obovate; latiseptate (flattened parallel to septum), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore at least 1.5 mm long, usually much longer, or absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Mexico.


Atelanthera

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively malpighiaceous (2-branched, sessile, medifixed); multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or China, or Pakistan, or Tajikistan, or Uzbekistan.


Athysanus

Heterodraba Greene.

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or stellate (star-shaped) and sessile, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals, or absent; white; obovate, or oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic, or ovate or obovate, or orbicular; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit two, or three to five, or six to ten, or eleven to 20; Seeds biseriate (in two rows), or aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Canada, or Mexico, or United States.


Aubrieta

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed, or a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet); obovate, or spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; filaments with a wing, tooth, or appendage; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or elliptic; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds biseriate (in two rows); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Iran, or Iraq, or Lebanon, or Syria, or Turkey, or Czech R, or Slovakia, or United States, or Ireland, or United Kingdom, or Crete, or Greece, or Bulgaria, or Yugoslavia, or Sicily, or France, or Albania, or Italy, or Romania, or Bosnia & Herz, or Croatia, or Macedonia.


Aurinia

Lepidotrichon Velen. & Bornm.

Annual or biennial, or perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) stellate (star-shaped) and sessile; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow; obovate; margin and apex entire. Sepals ascending; free. Stamens six; filaments with a wing, tooth, or appendage; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); oblong, or elliptic, or ovate or obovate; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds uniseriate (in one row), or biseriate (in two rows); wing present; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Armenia, or Turkey, or Moldova, or Ukraine, or Czech R, or Slovakia, or Slovenia, or Canada, or United States, or Greece, or Crete, or Albania, or Yugoslavia, or Italy, or Romania, or Bosnia & Herz, or Croatia, or Macedonia, or Austria, or France, or Germany, or Bulgaria, or Hungary, or Poland, or Russia.


Baimashania

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched; multicellular glands absent; not a weed. Stem leaves absent; divisions and margins entire or repand. Inflorescences solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals; white, or pink (or lavender); obovate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

China.


Ballantinia

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate, or orbicular; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic, or ovate or obovate; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Tasmania.


Barbarea

Barbamine A.P. Khokhr.; Campe Dulac.

Perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed, or a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; obovate, or oblong to elliptic; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing present, or absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or Armenia, or Azerbaijan, or Bhutan, or Chile, or Georgia, or India, or Iran, or Iraq, or Japan, or Kazakhstan, or Korea, or Kyrgyzstan, or Lebanon, or Mongolia, or Nepal, or Pakistan, or Russia, or Sri Lanka, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or Uzbekistan, or Yemen, or Argentina, or Estonia, or Latvia, or Lithuania, or Belarus, or Moldova, or Ukraine, or Czech R, or Slovakia, or Slovenia, or Canada, or Mexico, or United States, or Greenland, or Australia, or Tasmania, or Algeria, or Morocco, or Tunisia, or Ethiopia, or Kenya, or Uganda, or Tanzania, or Canary Islands, or Madeira, or Haiti, or New Zealand, or Taiwan, or South Africa, or Ireland, or United Kingdom, or Haiti, or Crete, or Greece, or Portugal, or Spain, or Albania, or Austria, or Belgium, or Bulgaria, or Bosnia & Herz, or Corsica, or Croatia, or Denmark, or Finland, or France, or Germany, or Netherlands, or Hungary, or Iceland, or Italy, or Macedonia, or Norway, or Poland, or Romania, or Sardinia, or Sicily, or Sweden, or Switzerland, or Yugoslavia.


Beringia

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50, or 51 to 100; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Canada, or Bosnia & Herz, or Belarus, or Bulgaria, or United States.


Berteroa

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or stellate (star-shaped) and sessile; multicellular glands absent; not a weed, or a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obcordate; margin and apex bifid, 3-lobed, pinnately lobed, or fimbriate. Sepals ascending, or spreading; free. Stamens six; filaments with a wing, tooth, or appendage; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); oblong, or elliptic, or ovate or obovate, or orbicular; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds biseriate (in two rows); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Armenia, or China, or Georgia, or Kazakhstan, or Kyrgyzstan, or Russia, or Tajikistan, or Turkey, or Uzbekistan, or Estonia, or Latvia, or Lithuania, or Belarus, or Moldova, or Ukraine, or Czech R, or Slovakia, or Slovenia, or Canada, or United States, or Ireland, or United Kingdom, or Greece, or Albania, or Austria, or Bulgaria, or Denmark, or Germany, or Switzerland, or Hungary, or Italy, or Yugoslavia, or Bosnia & Herz, or Croatia, or Macedonia, or Poland, or Romania, or Belgium, or Finland, or France, or Netherlands, or Norway, or Sweden.


Berteroella

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) stellate (star-shaped) and sessile; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; pink (or lavender), or purple (or violet); obovate, or oblong to elliptic; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

China, or Japan, or Korea.


Biscutella

Iondraba Medik.

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; oblong to elliptic, or spatulate to oblanceolate, or linear; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); didymous (spectacle shaped); angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages wings. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Cyprus, or Iran, or Iraq, or Israel, or Jordan, or Lebanon, or Saudi Arabia, or Syria, or Turkey, or Estonia, or Egypt, or Ukraine, or Czech R, or Slovakia, or Slovenia, or Algeria, or Libya, or Morocco, or Tunisia, or Canary Islands, or Madeira, or Crete, or Greece, or Portugal, or Spain, or Austria, or Belgium, or France, or Germany, or Switzerland, or Hungary, or Italy, or Yugoslavia, or Poland, or Romania, or Russia, or Balearic Islds, or Corsica, or Sardinia, or Sicily, or Albania, or Bosnia & Herz, or Croatia, or Macedonia.


Bivonaea

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; obovate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages wings. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Algeria, or Sardinia, or Sicily, or Tunisia.


Blennodia

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or stellate (star-shaped) and sessile, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); obovate, or oblong to elliptic; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything, or filaments with a wing, tooth, or appendage; filaments of median stamens free, or united at least along lower third; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50, or 51 to 100; Seeds biseriate (in two rows); wing present, or absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Australia.


Boechera

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple, or a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals; white, or pink (or lavender), or purple (or violet); obovate, or oblong to elliptic, or spatulate to oblanceolate, or linear; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted, or exserted well above sepals and petals. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50, or 51 to 100; Seeds uniseriate (in one row), or biseriate (in two rows); wing present, or absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Canada, or Mexico, or Russia, or United States, or Greenland.


Boreava

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; oblong to elliptic; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); globose (spherical), or ellipsoid; quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall woody or corky and thick; indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing lobes not decurrent; fruit appendages none, or wings. Ovules/seeds per ovary/fruit one; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Iran, or Israel, or Pakistan, or Turkey, or Syria, or Tunisia, or Spain.


Bornmuellera

Perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively malpighiaceous (2-branched, sessile, medifixed), or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate; margin and apex entire. Sepals ascending; free. Stamens six; filaments with a wing, tooth, or appendage; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); globose (spherical), or ovoid or obovoid, or ellipsoid; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall thinly papery, or neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit two, or three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Albania, or Greece, or Turkey, or Yugoslavia.


Borodinia

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; oblong to elliptic; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore at least 1.5 mm long, usually much longer, or absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Russia.


Botschantzevia

Perennial; shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or stellate (star-shaped) and sessile; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet); oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or oblong; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore at least 1.5 mm long, usually much longer, or absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds uniseriate (in one row); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Kazakhstan, or South Africa.


Brassica

Annual or biennial, or perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted, or exserted well above sepals and petals. Fruit a silique (fruit more than 3 times longer than broad); linear, or oblong; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore at least 1.5 mm long, usually much longer, or absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none, or a beaklike style or distal segment. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row), or biseriate (in two rows); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation segmented, or not segmented.

Afghanistan, or Armenia, or Azerbaijan, or Bahrain, or Bangladesh, or Bhutan, or Cambodia, or China, or Cyprus, or Georgia, or India, or Indonesia, or Iran, or Iraq, or Israel, or Japan, or Jordan, or Kazakhstan, or Korea, or Kuwait, or Kyrgyzstan, or Laos, or Lebanon, or Mongolia, or Myanmar, or Nepal, or Oman, or Pakistan, or Philippines, or Qatar, or Russia, or Saudi Arabia, or Sri Lanka, or Syria, or Tajikistan, or Thailand, or Turkmenistan, or Turkey, or United Arab Emir, or Uzbekistan, or Vietnam, or Yemen, or Argentina, or Bolivia, or Brazil, or Chile, or Colombia, or Ecuador, or Paraguay, or Peru, or Uruguay, or Venezuela, or Estonia, or Latvia, or Lithuania, or Belarus, or Moldova, or Egypt, or Ukraine, or Czech R, or Slovakia, or Slovenia, or Panama, or Costa Rica, or Guatemala, or Nicaragua, or Belize, or Honduras, or El Salvador, or Mexico, or United States, or Canada, or Bahamas, or Cuba, or Jamaica, or Haiti, or Dominican R, or Puerto Rico, or Lesser Antilles, or Greenland, or Australia, or Tasmania, or Morocco, or Libya, or Algeria, or Tunisia, or Kenya, or Tanzania, or Uganda, or Trinidad & Tobago, or Rwanda, or Zambia, or Zimbabwe, or Malawi, or Madagascar, or Eritrea, or Ethiopia, or Somalia, or Sudan, or Ivory Coast, or Sierra Leon, or Liberia, or Nigeria, or Congo Democratic R, or Cameron, or Papua NG, or Madeira, or Canary Islands, or Chad, or Sierra Leon, or Mali, or Mauritania, or New Caledonia, or New Zealand, or Cape Verde, or Taiwan, or Hawaii, or South Africa, or Namibia, or Botswana, or Ireland, or United Kingdom, or Other Pacific Islands, or Other Indian Islands, or Other Atlantic Islands, or Comoros, or Socotra, or Crete, or Greece, or Portugal, or Spain, or Bahamas, or Cuba, or Haiti, or Puerto Rico, or Jamaica, or Trinidad & Tobago, or French Guiana, or Surinam, or Sudan, or Djibouti, or Mozambique, or Zimbabwe, or Rwanda, or Niger, or Guyana, or Other Caribbean Islands, or Dominican R, or Angola, or Togo, or Senegal, or Burundi, or Central African R, or Congo, or Gabon, or Gambia, or Ghana, or Benin, or Guinea, or Mauritius, or Singapore, or Guinea Bissau, or Albania, or Austria, or Belgium, or Bosnia & Herz, or Bulgaria, or Corsica, or Croatia, or Denmark, or Finland, or France, or Germany, or Hungary, or Iceland, or Italy, or Macedonia, or Netherlands, or Norway, or Poland, or Romania, or Sardinia, or Sicily, or Sweden, or Switzerland, or Yugoslavia, or Balearic Islds, or Cameron, or Bourkina Fasso, or Equatorial Guinea.


Braya

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple, or a mixture of simple and branched; multicellular glands absent; not a weed. Stem leaves present, or absent; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are bracteate throughout or at least in lower half. Petals distinctly longer than sepals; white, or yellow, or pink (or lavender), or purple (or violet); obovate; margin and apex entire. Sepals ascending, or erect but not connivent; free, or united. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or oblong, or ovoid or obovoid; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row), or biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or Bhutan, or China, or India, or Kazakhstan, or Korea, or Kyrgyzstan, or Mongolia, or Nepal, or Pakistan, or Russia, or Tajikistan, or Canada, or United States, or Greenland, or Austria, or Italy, or Iceland, or Norway, or Sweden.


Brayopsis

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves absent; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals; white; obovate, or oblong to elliptic, or linear; margin and apex entire. Sepals ascending, or erect but not connivent, or spreading; free, or united. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or oblong; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina, or Bolivia, or Colombia, or Ecuador, or Peru.


Brossardia

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; pink (or lavender), or purple (or violet); obovate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovate or obovate, or orbicular; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall thinly papery; indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages wings. Ovules/seeds per ovary/fruit one, or two, or three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit reflexed (pendulous). Fruit segmentation not segmented.

Iran, or Iraq, or Turkey.


Bunias

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands present, or absent; not a weed, or a weed. Stem leaves present; base sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; obovate, or orbicular; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovoid or obovoid, or pyriform or obpyriform (pear-shaped); terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall woody or corky and thick; indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none, or wings, or surface verrucose. Ovules/seeds per ovary/fruit two, or three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon spiral; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Armenia, or Azerbaijan, or China, or Georgia, or Iran, or Iraq, or Kazakhstan, or Mongolia, or Russia, or Turkey, or Canada, or United States, or Albania, or Bulgaria, or Corsica, or Crete, or France, or Greece, or Switzerland, or Spain, or Italy, or Yugoslavia, or Portugal, or Sardinia, or Sicily, or Austria, or Czech R, or Germany, or Romania, or Hungary, or Poland, or Belgium, or United Kingdom, or Denmark, or Finland, or Netherlands, or Norway, or Sweden, or Estonia, or Latvia, or Lithuania, or Belarus, or Moldova, or Ukraine, or Slovakia, or Slovenia, or Algeria, or Tunisia, or Ireland, or United Kingdom, or Spain.


Cakile

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals, or reduced and smaller than sepals, or absent; white, or pink (or lavender), or purple (or violet); obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); lanceolate or oblanceolate, or oblong, or ovoid or obovoid; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick; breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit one, or two, or three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation segmented.

Argentina, or Cyprus, or Georgia, or Iran, or Iraq, or Israel, or Japan, or Kuwait, or Lebanon, or Saudi Arabia, or Syria, or Turkey, or Uruguay, or Estonia, or Latvia, or Lithuania, or Belarus, or Egypt, or Algeria, or Belgium, or United Kingdom, or United States, or France, or Germany, or Greece, or Crete, or Ireland, or Italy, or Morocco, or Libya, or Tunisia, or Portugal, or Spain, or Yugoslavia, or Australia, or Canada, or Mexico, or New Caledonia, or Denmark, or Finland, or Poland, or Sweden, or Bulgaria, or Romania, or Russia, or Lesser Antilles, or Cuba, or Colombia, or Dominican R, or Haiti, or Honduras, or Jamaica, or Panama, or Venezuela, or Belize, or Tasmania, or New Zealand, or Ukraine, or Czech R, or Slovakia, or Slovenia, or Puerto Rico, or Bahamas, or Armenia, or Canary Islands, or Madeira, or Cape Verde, or Other Pacific Islands, or Other Indian Islands, or Other Atlantic Islands.


Calepina

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed, or a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovoid or obovoid, or ellipsoid, or pyriform or obpyriform (pear-shaped); terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick; indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit one; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers zygomorphic (two petals and/or sepals much larger than other two); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Armenia, or Azerbaijan, or Cyprus, or Georgia, or Iran, or Iraq, or Israel, or Jordan, or Lebanon, or Syria, or Turkmenistan, or Turkey, or Ukraine, or Slovenia, or United States, or Algeria, or Libya, or Morocco, or Crete, or Greece, or Portugal, or Spain, or Albania, or Belgium, or Bulgaria, or Corsica, or France, or Germany, or Austria, or Switzerland, or Netherlands, or Hungary, or Italy, or Yugoslavia, or Romania, or Russia, or Sardinia, or Sicily.


Calymmatium

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow; oblong to elliptic; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or oblong, or ellipsoid; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or Tajikistan.


Camelina

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed, or a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow; obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); oblong, or pyriform or obpyriform (pear-shaped); terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick, or neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none, or wings. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds biseriate (in two rows); wing absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or Argentina, or Armenia, or Azerbaijan, or China, or Cyprus, or Georgia, or India, or Iran, or Iraq, or Ireland, or Japan, or Jordan, or Kazakhstan, or Korea, or Kyrgyzstan, or Lebanon, or Mongolia, or Pakistan, or Russia, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or Uzbekistan, or Estonia, or Latvia, or Lithuania, or Belarus, or Moldova, or Egypt, or Ukraine, or Czech R, or Slovakia, or Slovenia, or Canada, or Australia, or Mexico, or United States, or Algeria, or Libya, or Morocco, or Tunisia, or Kenya, or Uganda, or New Zealand, or Ireland, or United Kingdom, or Crete, or Greece, or Portugal, or Spain, or Albania, or Austria, or Belgium, or Bosnia & Herz, or Bulgaria, or Corsica, or Croatia, or Denmark, or France, or Germany, or Hungary, or Iceland, or Italy, or Macedonia, or Norway, or Poland, or Romania, or Sardinia, or Sicily, or Sweden, or Yugoslavia.


Camelinopsis

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; oblong to elliptic; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovate or obovate, or ovoid or obovoid; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit reflexed (pendulous). Fruit segmentation not segmented.

Iran, or Iraq.


Capsella

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or stellate (star-shaped) and sessile; multicellular glands absent; not a weed, or a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals, or reduced and smaller than sepals, or absent; white; obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); triangular, or cordate or obcordate; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina, or Bolivia, or Ecuador, or Peru, or Afghanistan, or Armenia, or Azerbaijan, or Bhutan, or China, or Cyprus, or Georgia, or India, or Indonesia, or Iran, or Iraq, or Israel, or Japan, or Jordan, or Kazakhstan, or Korea, or Lebanon, or Mongolia, or Nepal, or Oman, or Pakistan, or Philippines, or Russia, or Saudi Arabia, or Sri Lanka, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or United Arab Emir, or Uzbekistan, or Vietnam, or Yemen, or Estonia, or Latvia, or Lithuania, or Belarus, or Kyrgyzstan, or Moldova, or Egypt, or Ukraine, or Czech R, or Slovakia, or Slovenia, or Canada, or Mexico, or United States, or Greenland, or Australia, or Tasmania, or Algeria, or Libya, or Morocco, or Tunisia, or Kenya, or Tanzania, or Uganda, or Zimbabwe, or Zambia, or Burundi, or Congo Democratic R, or Ethiopia, or Sudan, or New Zealand, or Madeira, or Canary Islands, or Taiwan, or Hawaii, or Lesotho, or South Africa, or Colombia, or Brazil, or Ireland, or United Kingdom, or Other Pacific Islands, or Other Indian Islands, or Other Atlantic Islands, or Socotra, or Crete, or Greece, or Portugal, or Spain, or Cuba, or Haiti, or Jamaica, or Djibouti, or Zimbabwe, or Rwanda, or Dominican R, or Congo.


Cardamine

Dentaria L.; Loxostemon Hook. f. & Thomson.

Annual or biennial, or perennial; herbs; terrestrial, or aquatic; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed, or a weed. Stem leaves present, or absent; base petiolate, or sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid), or palmately lobed, or 1–3-pinnatisect, or trifoliolate, or pinnately compound, or palmately compound. Inflorescences racemes that are bracteate throughout or at least in lower half, or racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate, or solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals, or subequalling sepals, or reduced and smaller than sepals, or absent; white, or pink (or lavender), or purple (or violet); obovate, or oblong to elliptic, or orbicular, or spatulate to oblanceolate, or obcordate; margin and apex entire. Sepals ascending, or erect but not connivent, or spreading; free. Stamens six; lower part of filaments and petal claws without anything, or filaments with a wing, tooth, or appendage; filaments of median stamens free; stamens included within calyx or partially exserted, or exserted well above sepals and petals. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing present, or absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate, or opposite, or whorled. Flowers actinomorphic (all petals equal), or zygomorphic (two petals and/or sepals much larger than other two); All flowers bisexual (with stamens and pistils), or At least some flowers unisexual. Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Argentina, or Bolivia, or Chile, or Colombia, or Ecuador, or Peru, or Venezuela, or Afghanistan, or Armenia, or Azerbaijan, or Bangladesh, or Bhutan, or China, or Cyprus, or Georgia, or India, or Indonesia, or Iran, or Iraq, or Israel, or Japan, or Jordan, or Kazakhstan, or Korea, or Kyrgyzstan, or Laos, or Lebanon, or Malaysia, or Mongolia, or Myanmar, or Nepal, or Pakistan, or Papua NG, or Philippines, or Russia, or Saudi Arabia, or Sri Lanka, or Syria, or Tajikistan, or Thailand, or Turkmenistan, or Turkey, or Uzbekistan, or Yemen, or Estonia, or Latvia, or Lithuania, or Belarus, or Moldova, or Ukraine, or Czech R, or Slovakia, or Slovenia, or Lesser Antilles, or Canada, or United States, or Mexico, or Belize, or Costa Rica, or Guatemala, or Nicaragua, or Honduras, or Panama, or El Salvador, or Greenland, or Australia, or Tasmania, or Tunisia, or Morocco, or Libya, or Algeria, or Kenya, or Tanzania, or Uganda, or Zambia, or Zimbabwe, or Taiwan, or Comoros, or Madagascar, or Nigeria, or Cameron, or Congo Democratic R, or Burundi, or Rwanda, or Eritrea, or Ethiopia, or Sudan, or Somalia, or New Caledonia, or Singapore, or Canary Islands, or New Zealand, or Madeira, or Canary Islands, or Taiwan, or Hawaii, or South Africa, or Brazil, or Uruguay, or Paraguay, or Ireland, or United Kingdom, or Other Pacific Islands, or Other Indian Islands, or Other Atlantic Islands, or Crete, or Greece, or Portugal, or Spain, or Bahamas, or Cuba, or Haiti, or Puerto Rico, or Jamaica, or Djibouti, or Zimbabwe, or Rwanda, or Albania, or Austria, or Belgium, or Bosnia & Herz, or Bulgaria, or Corsica, or Croatia, or Denmark, or Finland, or France, or Balearic Islds, or Germany, or Netherlands, or Hungary, or Iceland, or Italy, or Macedonia, or Norway, or Poland, or Romania, or Sardinia, or Sicily, or Sweden, or Switzerland, or Yugoslavia, or Angola, or Congo, or Gabon, or Equatorial Guinea.


Carinavalva

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free, or united at least along lower third; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic, or ovate or obovate; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit more than 100; Seeds biseriate (in two rows); wing present; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Australia.


Carrichtera

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate; divisions and margins 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; obovate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovoid or obovoid, or ellipsoid; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages wings, or a beaklike style or distal segment. Ovules/seeds per ovary/fruit three to five, or six to ten; Seeds biseriate (in two rows); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit reflexed (pendulous). Fruit segmentation segmented.

Cyprus, or Iran, or Iraq, or Israel, or Japan, or Jordan, or Kuwait, or Lebanon, or Saudi Arabia, or Syria, or Egypt, or Australia, or Algeria, or Libya, or Morocco, or Tunisia, or Canary Islands, or New Zealand, or United Kingdom, or Crete, or Greece, or Spain, or Balearic Islds, or Sardinia, or Sicily.


Catadysia

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves absent; divisions and margins dentate, serrate, incised, crenate, sinuate. Inflorescences solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals; white; obovate; margin and apex entire. Sepals ascending; united. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); oblong; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Peru.


Catenulina

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively malpighiaceous (2-branched, sessile, medifixed); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals reduced and smaller than sepals; white; oblong to elliptic; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds uniseriate (in one row); wing present; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Tajikistan.


Catolobus

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals; white, or yellow, or pink (or lavender); oblong to elliptic, or spatulate to oblanceolate, or linear; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50, or 51 to 100, or more than 100; Seeds uniseriate (in one row), or biseriate (in two rows); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit reflexed (pendulous). Fruit segmentation not segmented.

China, or Japan, or Kazakhstan, or Korea, or Mongolia, or Russia.


Ceratocnemum

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); lanceolate or oblanceolate, or oblong; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick; breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages horns, or a beaklike style or distal segment. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit segmentation segmented.

Morocco.


Chalcanthus

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; purple (or violet); obovate, or oblong to elliptic; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50, or 51 to 100; Seeds uniseriate (in one row); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or Iran, or Kazakhstan, or Kyrgyzstan, or Tajikistan, or Turkmenistan, or Uzbekistan.


Chamira

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; oblong to elliptic; margin and apex entire. Sepals erect and connivent; free. Stamens six; filaments with a wing, tooth, or appendage; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); linear, or oblong; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten; Seeds uniseriate (in one row); wing absent; cotyledon of other types; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate, or opposite. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

South Africa.


Chartoloma

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; oblong to elliptic; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall thinly papery; indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing lobes not decurrent; fruit appendages wings. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit reflexed (pendulous). Fruit segmentation not segmented.

Turkmenistan, or Uzbekistan.


Chaunanthus

Perennial; shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple, or a mixture of simple and branched; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50, or 51 to 100; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Mexico.


Chilocardamum

Dimitria Ravenna.

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present, or absent; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything, or petal claw and/or filament with hairs or papillae; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50; Seeds uniseriate (in one row); wing present, or absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina.


Chlorocrambe

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens exserted well above sepals and petals. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall thinly papery; dehiscent (readily releasing seeds); gynophore at least 1.5 mm long, usually much longer, or absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

United States.


Chorispora

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands present; not a weed, or a weed. Stem leaves present, or absent; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate, or solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals; white, or yellow, or pink (or lavender), or purple (or violet); obovate, or oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick; breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages none, or a beaklike style or distal segment. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation segmented, or not segmented.

Afghanistan, or Argentina, or Armenia, or Azerbaijan, or China, or Georgia, or India, or Iran, or Iraq, or Israel, or Japan, or Jordan, or Kazakhstan, or Korea, or Kyrgyzstan, or Lebanon, or Mongolia, or Pakistan, or Russia, or Saudi Arabia, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or Uzbekistan, or Estonia, or Belarus, or Moldova, or Ukraine, or Czech R, or Slovakia, or Slovenia, or Canada, or United States, or Algeria, or Greece, or Bulgaria, or Yugoslavia, or Romania.


Christolea

Koelzia Rech. f.

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or lanceolate or oblanceolate; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or China, or India, or Nepal, or Pakistan, or Tajikistan.


Chrysochamela

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals; yellow; obovate; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything, or filaments with a wing, tooth, or appendage; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); oblong, or elliptic, or ovate or obovate; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Syria, or Turkey, or Russia.


Cithareloma

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); obovate, or oblong to elliptic; margin and apex entire. Sepals erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); oblong, or elliptic; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds uniseriate (in one row); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or China, or Iran, or Kazakhstan, or Kyrgyzstan, or Pakistan, or Tajikistan, or Turkmenistan, or Uzbekistan.


Clastopus

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow, or purple (or violet); obovate; margin and apex entire. Sepals erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything, or filaments with a wing, tooth, or appendage; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovate or obovate, or orbicular; latiseptate (flattened parallel to septum); fruit (or segments) wall woody or corky and thick, or neither woody or papery (normal); indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Iran, or Iraq, or Turkey.


Clausia

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands present; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; purple (or violet); obovate; margin and apex entire. Sepals erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50; Seeds uniseriate (in one row); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or China, or Kazakhstan, or Korea, or Mongolia, or Russia, or Kyrgyzstan, or Ukraine.


Clypeola

Pseudoanastatica Grossh.

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) stellate (star-shaped) and sessile, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; filaments with a wing, tooth, or appendage; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); orbicular; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none, or wings, or glochidiate trichomes. Ovules/seeds per ovary/fruit one; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit reflexed (pendulous). Fruit segmentation not segmented.

Afghanistan, or Armenia, or Azerbaijan, or Cyprus, or Georgia, or Iran, or Iraq, or Israel, or Jordan, or Kazakhstan, or Kyrgyzstan, or Lebanon, or Oman, or Panama, or Saudi Arabia, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or Uzbekistan, or Egypt, or Libya, or Tunisia, or Algeria, or Morocco, or Albania, or Bulgaria, or Corsica, or Crete, or France, or Greece, or Switzerland, or Spain, or Italy, or Yugoslavia, or Romania, or Russia, or Sardinia, or Sicily, or Ukraine, or Balearic Islds.


Cochlearia

Cochleariopsis A. & D. Löve; Glaucocochlearia (O.E. Schulz) Pobed.; Ionopsidium Rchb.

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed, or a weed. Stem leaves present, or absent; base petiolate, or sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate, or solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals; white, or yellow, or pink (or lavender), or purple (or violet); obovate, or oblong to elliptic; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); oblong, or elliptic, or ovate or obovate, or orbicular, or globose (spherical); angustiseptate (flattened at a right angle to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none, or wings. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row), or biseriate (in two rows); wing absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Falkland Isl, or Armenia, or Japan, or Russia, or Turkey, or Estonia, or Ukraine, or Czech R, or Slovakia, or Canada, or United States, or Greenland, or Morocco, or Ireland, or United Kingdom, or Portugal, or Spain, or Belgium, or Denmark, or Finland, or France, or Greece, or Netherlands, or Norway, or Austria, or Other Atlantic Islands, or France, or Germany, or Iceland, or Poland, or Romania, or Sweden, or Italy, or Algeria, or Tunisia, or Sicily.


Coelophragmus

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or auriculate, amplexicaul, or sagittate; divisions and margins pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit 51 to 100, or more than 100; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Mexico, or United States.


Coincya

Brassicella Fourr. Ex O.E. Schulz; Hutera Porta; Rhynchosinapis Hayek.

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow; obovate; margin and apex entire. Sepals erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or oblong; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick, or neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none, or a beaklike style or distal segment. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50, or 51 to 100, or more than 100; Seeds uniseriate (in one row); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation segmented.

Japan, or United States, or Morocco, or United Kingdom, or Greece, or Portugal, or Spain, or France, or Italy, or Belgium, or Corsica, or France, or Germany, or Sardinia, or Netherlands.


Coluteocarpus

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; oblong to elliptic; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovoid or obovoid, or ellipsoid; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall thinly papery; indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds uniseriate (in one row), or aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit reflexed (pendulous). Fruit segmentation not segmented.

Armenia, or Azerbaijan, or Georgia, or Iran, or Iraq, or Lebanon, or Syria, or Turkey.


Conringia

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed, or a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow; orbicular, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes decurrent (connivent); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or Armenia, or Azerbaijan, or Georgia, or China, or Cyprus, or Iran, or Iraq, or Israel, or Japan, or Jordan, or Kazakhstan, or Kyrgyzstan, or Lebanon, or Mongolia, or Pakistan, or Russia, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or Uzbekistan, or Estonia, or Lithuania, or Belarus, or Egypt, or Ukraine, or Czech R, or Slovakia, or Slovenia, or Canada, or Mexico, or United States, or Australia, or Libya, or Algeria, or Morocco, or Tunisia, or Ireland, or United Kingdom, or Greece, or Spain, or Bahamas, or Albania, or Austria, or Balearic Islds, or Bosnia & Herz, or Bulgaria, or Croatia, or Denmark, or France, or Germany, or Hungary, or Iceland, or Italy, or Macedonia, or Norway, or Poland, or Romania, or Sardinia, or Netherlands, or Switzerland, or Yugoslavia.


Cordylocarpus

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; obovate, or orbicular; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick; breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages wings. Ovules/seeds per ovary/fruit two, or three to five; Seeds uniseriate (in one row); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit segmentation segmented.

Algeria, or Morocco, or Israel.


Crambe

Perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow; obovate, or oblong to elliptic; margin and apex entire. Sepals ascending; free. Stamens six; filaments with a wing, tooth, or appendage; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); globose (spherical); terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick; breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation segmented.

Afghanistan, or Armenia, or Azerbaijan, or China, or Cyprus, or Georgia, or India, or Iran, or Iraq, or Israel, or Jordan, or Kazakhstan, or Kyrgyzstan, or Lebanon, or Pakistan, or Russia, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or Uzbekistan, or Estonia, or Lithuania, or Moldova, or Ukraine, or Czech R, or Slovakia, or Australia, or Algeria, or Morocco, or Libya, or Kenya, or Tanzania, or Uganda, or Rwanda, or Congo Democratic R, or Ethiopia, or Canary Islands, or Chad, or Madeira, or Eritrea, or South Africa, or Ireland, or United Kingdom, or Greece, or Portugal, or Spain, or Rwanda, or Austria, or Belgium, or Bulgaria, or Denmark, or Finland, or France, or Germany, or Netherlands, or Norway, or Romania, or Sweden, or Hungary, or Italy, or Yugoslavia, or Albania, or Bosnia & Herz, or Croatia, or Macedonia, or Corsica, or Sardinia, or Sicily.


Crambella

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; purple (or violet); obovate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); globose (spherical); terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick; breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit two, or three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation segmented.

Morocco.


Cremolobus

Loxoptera O.E. Schulz; Urbanodoxa Muschl.

Annual or biennial, or perennial; herbs, or shrubs or subshrubs, or lianas; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple, or stellate (star-shaped) and sessile; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow; obovate, or oblong to elliptic; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); didymous (spectacle shaped); angustiseptate (flattened at a right angle to septum); fruit (or segments) wall woody or corky and thick; breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore at least 1.5 mm long, usually much longer, or absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none, or wings. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate, or opposite. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina, or Bolivia, or Chile, or Colombia, or Ecuador, or Peru.


Crucihimalaya

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like), or stellate and stalked; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are bracteate throughout or at least in lower half, or racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50, or 51 to 100, or more than 100; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or Bhutan, or China, or India, or Iran, or Kazakhstan, or Kyrgyzstan, or Mongolia, or Nepal, or Pakistan, or Russia, or Saudi Arabia, or Tajikistan, or Turkmenistan, or Uzbekistan, or Egypt.


Cryptospora

Trichotochiton Kom.

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick; breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or China, or Iran, or Kazakhstan, or Kyrgyzstan, or Tajikistan, or Turkmenistan, or Uzbekistan.


Cuphonotus

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow; spatulate to oblanceolate, or linear; margin and apex entire. Sepals spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic, or orbicular; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds uniseriate (in one row); wing present; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Australia.


Cusickeilla

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present, or absent; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow; oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovoid or obovoid, or ellipsoid; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

United States.


Cycloptychis

Perennial; shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or purple (or violet); oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic, or ovate or obovate; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none, or a beaklike style or distal segment. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon spiral; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

South Africa.


Cymatocarpus

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; obovate, or orbicular; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Afghanistan, or Armenia, or Azerbaijan, or Iran, or Tajikistan, or Turkmenistan, or Uzbekistan.


Cyphocardamum

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present, or absent; base sessile but not auriculate or amplexicaul; divisions and margins dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate, or oblong to elliptic; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovate or obovate; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan.


Dactylocardamum

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals; white; spatulate to oblanceolate, or linear; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovoid or obovoid; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit one, or two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Peru.


Degenia

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) stellate (star-shaped) and sessile; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; spatulate to oblanceolate; margin and apex entire. Sepals erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic, or ovate or obovate, or orbicular; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall thinly papery, or neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five; Seeds biseriate (in two rows); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Croatia.


Delpinophytum

Delpinoella Speg.

Perennial; shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves absent; divisions and margins entire or repand. Inflorescences solitary flowers borne on pedicels originating from basal rosette. Petals subequalling sepals; white; oblong to elliptic; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); didymous (spectacle shaped); angustiseptate (flattened at a right angle to septum); fruit (or segments) wall woody or corky and thick; breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina.


Descurainia

Sophia Adans.; Hugenenia Rchb.; Robeschia Hoschst. ex O.E. Schulz.

Annual or biennial, or perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) at least some dendritic (tree-like); multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate; divisions and margins 1–3-pinnatisect. Inflorescences racemes that are bracteate throughout or at least in lower half, or racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals; white, or yellow; obovate, or oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or spreading, or reflexed; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or oblong, or elliptic; terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20, or 21 to 50, or 51 to 100; Seeds uniseriate (in one row), or biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina, or Bolivia, or Chile, or Ecuador, or Peru, or Afghanistan, or Armenia, or Azerbaijan, or Bhutan, or China, or Georgia, or India, or Iran, or Iraq, or Israel, or Japan, or Jordan, or Kazakhstan, or Kyrgyzstan, or Korea, or Kuwait, or Lebanon, or Mongolia, or Nepal, or Pakistan, or Russia, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or Uzbekistan, or Estonia, or Latvia, or Lithuania, or Belarus, or Moldova, or Egypt, or Ukraine, or Czech R, or Slovakia, or Slovenia, or Canada, or United States, or Mexico, or Guatemala, or Greenland, or Australia, or Algeria, or Morocco, or Canary Islands, or New Zealand, or Hawaii, or South Africa, or Ireland, or United Kingdom, or Crete, or Greece, or Portugal, or Spain, or Albania, or Austria, or Belgium, or Bosnia & Herz, or Bulgaria, or Corsica, or Croatia, or Denmark, or Finland, or France, or Germany, or Netherlands, or Hungary, or Iceland, or Italy, or Macedonia, or Norway, or Poland, or Romania, or Sardinia, or Sicily, or Sweden, or Switzerland, or Yugoslavia, or Balearic Islds.


Desideria

Ermaniopsis H. Hara; Oreoblastus Suslova.

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched; multicellular glands absent; not a weed. Stem leaves present, or absent; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or palmately lobed. Inflorescences racemes that are bracteate throughout or at least in lower half, or racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate, or solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals; white, or purple (or violet); obovate, or oblong to elliptic; margin and apex entire. Sepals erect but not connivent, or erect and connivent; free, or united. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50, or 51 to 100; Seeds uniseriate (in one row), or biseriate (in two rows); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or China, or India, or Kazakhstan, or Kyrgyzstan, or Nepal, or Pakistan, or Tajikistan, or Turkmenistan.


Diceratella

Perennial; shrubs or subshrubs; terrestrial; thorns and spines present, or absent; hairs (trichomes) stellate (star-shaped) and sessile, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet); obovate; margin and apex entire. Sepals erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or oblong; terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages horns. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Iran, or Pakistan, or Yemen, or Algeria, or Kenya, or Ethiopia, or Socotra, or Somalia, or Sudan, or Djibouti, or Egypt, or Kenya, or Ethiopia, or Somalia, or Socotra, or Sudan, or Djibouti.


Dichasianthus

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; pink (or lavender), or purple (or violet); oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or Kazakhstan, or Tajikistan, or Turkmenistan.


Dictyophragmus

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are bracteate throughout or at least in lower half, or racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate, or spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything, or petal claw and/or filament with hairs or papillae; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or lanceolate or oblanceolate; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina, or Peru.


Didesmus

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate, or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); obovate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); lanceolate or oblanceolate, or ovoid or obovoid; terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall woody or corky and thick; breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation segmented.

Cyprus, or Crete, or Greece, or Egypt, or Libya, or Tunisia, or Algeria.


Didymophysa

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or palmately lobed. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet); obovate, or orbicular; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); didymous (spectacle shaped); terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall thinly papery; indehiscent (seeds remain inside whole fruit), or breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or Armenia, or Azerbaijan, or Iran, or Iraq, or Kyrgyzstan, or Pakistan, or Tajikistan, or Turkey, or Uzbekistan.


Dielsiocharis

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; obovate, or oblong to elliptic; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); oblong; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20, or 21 to 50; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Iran, or Tajikistan.


Dilophia

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are bracteate throughout or at least in lower half, or racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); obovate, or obcordate; margin and apex entire. Sepals spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); cordate or obcordate; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed); fruit appendages none, or crests. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds uniseriate (in one row), or aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Bhutan, or China, or India, or Kyrgyzstan, or Nepal, or Pakistan, or Tajikistan.


Dimorphocarpa

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); obovate; margin and apex entire. Sepals spreading, or reflexed; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); didymous (spectacle shaped); angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Mexico, or United States.


Diplotaxis

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow, or pink (or lavender), or purple (or violet); obovate, or oblong to elliptic, or orbicular; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore at least 1.5 mm long, usually much longer, or absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50, or 51 to 100, or more than 100; Seeds biseriate (in two rows); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation segmented, or not segmented.

Argentina, or Ecuador, or Peru, or Afghanistan, or Azerbaijan, or Bahrain, or China, or Cyprus, or Georgia, or India, or Iran, or Iraq, or Israel, or Japan, or Jordan, or Kuwait, or Lebanon, or Nepal, or Oman, or Pakistan, or Saudi Arabia, or Syria, or Turkey, or United Arab Emir, or Yemen, or Estonia, or Latvia, or Lithuania, or Kazakhstan, or Belarus, or Moldova, or Egypt, or Ukraine, or Czech R, or Slovakia, or Slovenia, or Canada, or Mexico, or United States, or Costa Rica, or Australia, or Tasmania, or Algeria, or Libya, or Morocco, or Tunisia, or Somalia, or Mauritania, or Cape Verde, or Chad, or New Caledonia, or New Zealand, or Madeira, or South Africa, or Brazil, or Ireland, or United Kingdom, or Other Atlantic Islands, or Bahamas, or Cuba, or Haiti, or Dominican R, or Crete, or Greece, or Portugal, or Spain, or Balearic Islds, or Sicily, or Albania, or Austria, or Belgium, or Bosnia & Herz, or Bulgaria, or Corsica, or Croatia, or Denmark, or Finland, or France, or Germany, or Switzerland, or Netherlands, or Hungary, or Italy, or Macedonia, or Norway, or Poland, or Romania, or Russia, or Sardinia, or Sicily, or Sweden, or Yugoslavia.


Dipoma

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or exclusively malpighiaceous (2-branched, sessile, medifixed); multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are bracteate throughout or at least in lower half. Petals distinctly longer than sepals; white; orbicular, or obcordate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovate or obovate; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none, or wings. Ovules/seeds per ovary/fruit three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

China.


Diptychocarpus

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands present; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); linear; margin and apex entire. Sepals erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick, or neither woody or papery (normal); dehiscent (readily releasing seeds), or breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or Armenia, or Azerbaijan, or China, or Iran, or Kazakhstan, or Kyrgyzstan, or Pakistan, or Tajikistan, or Turkmenistan, or Uzbekistan.


Dithyrea

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); spatulate to oblanceolate; margin and apex entire. Sepals erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); didymous (spectacle shaped); angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Mexico, or United States.


Dontostemon

Dimorphostemon Kitag.; Alaida F. Dvorak.

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands present, or absent; not a weed, or a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); obovate, or spatulate to oblanceolate, or obcordate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything, or filaments with a wing, tooth, or appendage; filaments of median stamens free, or united at least along lower third; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes decurrent (connivent); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50, or 51 to 100; Seeds uniseriate (in one row); wing present, or absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Bhutan, or China, or India, or Japan, or Kazakhstan, or Korea, or Mongolia, or Nepal, or Pakistan, or Russia, or Tajikistan, or Kyrgyzstan.


Douepea

Dolichorhynchus Hedge & Kit Tan.

Perennial; shrubs or subshrubs; terrestrial; thorns and spines present, or absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); linear; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50; Seeds uniseriate (in one row); wing present, or absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation segmented.

Pakistan, or Saudi Arabia.


Draba

Drabopsis K. Koch; Erophila DC.; Coelonema Maxim.; Schivereckia Andrz. ex DC.

Annual or biennial, or perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or exclusively malpighiaceous (2-branched, sessile, medifixed), or stellate (star-shaped) and sessile, or at least some dendritic (tree-like); multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are bracteate throughout or at least in lower half, or racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals, or reduced and smaller than sepals, or absent; white, or yellow, or pink (or lavender), or purple (or violet), or orange; obovate, or oblong to elliptic, or spatulate to oblanceolate, or obcordate; margin and apex entire, or bifid, 3-lobed, pinnately lobed, or fimbriate. Sepals ascending, or erect but not connivent, or spreading; free. Stamens six, or four; lower part of filaments and petal claws without anything, or filaments with a wing, tooth, or appendage; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or lanceolate or oblanceolate, or oblong, or elliptic, or ovate or obovate, or orbicular, or globose (spherical), or ovoid or obovoid, or ellipsoid; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20, or 21 to 50, or 51 to 100; Seeds biseriate (in two rows); wing present, or absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two, or three to six. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Argentina, or Bolivia, or Chile, or Colombia, or Ecuador, or Peru, or Uruguay, or Venezuela, or Falkland Isl, or Afghanistan, or Armenia, or Azerbaijan, or Bhutan, or China, or Cyprus, or Georgia, or India, or Iran, or Iraq, or Israel, or Japan, or Jordan, or Kazakhstan, or Korea, or Kyrgyzstan, or Lebanon, or Mongolia, or Nepal, or Pakistan, or Russia, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or Uzbekistan, or Yemen, or Estonia, or Latvia, or Lithuania, or Belarus, or Moldova, or Ukraine, or Czech R, or Slovakia, or Slovenia, or Canada, or Costa Rica, or Guatemala, or Mexico, or United States, or Greenland, or Tasmania, or Australia, or Algeria, or Libya, or Morocco, or Tunisia, or Ethiopia, or New Zealand, or Madeira, or Taiwan, or Ireland, or United Kingdom, or Crete, or Greece, or Portugal, or Spain, or Albania, or Austria, or Belgium, or Bosnia & Herz, or Bulgaria, or Corsica, or Croatia, or Denmark, or Finland, or France, or Germany, or Hungary, or Iceland, or Italy, or Macedonia, or Norway, or Poland, or Romania, or Sardinia, or Sicily, or Sweden, or Switzerland, or Yugoslavia.


Drabastrum

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); obovate, or oblong to elliptic, or orbicular; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic, or ovate or obovate; terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds biseriate (in two rows); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Australia.


Dryopetalon

Rollinsia Al-Shehbaz.

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet); obovate, or oblong to elliptic, or spatulate to oblanceolate; margin and apex entire, or bifid, 3-lobed, pinnately lobed, or fimbriate. Sepals ascending, or erect but not connivent, or spreading; free. Stamens six; lower part of filaments and petal claws without anything, or petal claw and/or filament with hairs or papillae; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore at least 1.5 mm long, usually much longer, or absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none, or crests. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Mexico, or United States.


Eigia

Microstigmatella Rauschert; Stigmatella Eig.

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present, or absent; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; purple (or violet); obovate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or oblong; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Israel, or Saudi Arabia.


Elburzia

Perennial; shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins palmately lobed. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; orbicular; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); ellipsoid; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Iran.


Enarthrocapus

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are bracteate throughout or at least in lower half. Petals distinctly longer than sepals; yellow; obovate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick; breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation segmented.

Cyprus, or Israel, or Jordan, or Lebanon, or Pakistan, or Saudi Arabia, or Syria, or Turkey, or Egypt, or Libya, or Tunisia, or Algeria, or Morocco, or Crete, or Greece, or Bulgaria.


Englerocharis

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves absent; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); oblong; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Peru.


Eremobium

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) stellate (star-shaped) and sessile; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are bracteate throughout or at least in lower half. Petals distinctly longer than sepals; pink (or lavender), or purple (or violet); oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Iran, or Iraq, or Israel, or Jordan, or Oman, or Pakistan, or Qatar, or Saudi Arabia, or United Arab Emir, or Yemen, or Algeria, or Egypt, or Libya, or Tunisia, or Morocco, or Mali, or Mauritania, or Niger.


Eremoblastus

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) stellate (star-shaped) and sessile, or at least some dendritic (tree-like); multicellular glands present; not a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; oblong to elliptic; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50; Seeds uniseriate (in one row); wing present; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Kazakhstan, or Russia.


Eremodraba

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; petal claw and/or filament with hairs or papillae; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); lanceolate or oblanceolate, or oblong; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Chile, or Peru.


Eremophyton

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; purple (or violet); obovate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); oblong, or ellipsoid; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick; breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit two, or three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation segmented.

Algeria, or Libya, or Morocco.


Eruca

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; obovate; margin and apex entire. Sepals erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or oblong, or ellipsoid; terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages a beaklike style or distal segment. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50; Seeds biseriate (in two rows); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation segmented.

Argentina, or Bolivia, or Ecuador, or Afghanistan, or Armenia, or Azerbaijan, or Bahrain, or China, or Cyprus, or Georgia, or India, or Iran, or Iraq, or Israel, or Japan, or Jordan, or Kazakhstan, or Kuwait, or Kyrgyzstan, or Lebanon, or Mongolia, or Oman, or Pakistan, or Qatar, or Russia, or Saudi Arabia, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or United Arab Emir, or Uzbekistan, or Yemen, or Estonia, or Latvia, or Lithuania, or Belarus, or Moldova, or Egypt, or Ukraine, or Czech R, or Slovenia, or Mexico, or Canada, or United States, or Australia, or Algeria, or Libya, or Morocco, or Tunisia, or Mauritania, or Sudan, or Ethiopia, or Eritrea, or Somalia, or Kenya, or Zambia, or Zimbabwe, or Canary Islands, or Madeira, or New Zealand, or South Africa, or Ireland, or United Kingdom, or Sudan, or Zimbabwe, or Crete, or Greece, or Portugal, or Spain, or Albania, or Austria, or Balearic Islds, or Bosnia & Herz, or Bulgaria, or Corsica, or Croatia, or France, or Switzerland, or Italy, or Yugoslavia, or Macedonia, or Romania, or Germany, or Norway, or Poland.


Erucaria

Hussonia Boiss.; Reboudia Coss. & Durieu ex Coss.

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet); obovate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or elliptic; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick, or neither woody or papery (normal); dehiscent (readily releasing seeds), or breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon incumbent, or conduplicate, or diplecolobal, or spiral; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation segmented.

Bahrain, or Cyprus, or Iran, or Iraq, or Israel, or Jordan, or Kuwait, or Lebanon, or Oman, or Pakistan, or Saudi Arabia, or Syria, or Turkey, or United Arab Emir, or Algeria, or Egypt, or Lithuania, or Tunisia, or Morocco, or Crete, or Greece, or Spain, or Libya, or Morocco, or Tunisia, or Crete, or Greece, or Spain.


Erucastrum

Hirschfeldia Moench.

Annual or biennial, or perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are bracteate throughout or at least in lower half, or racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow; obovate, or oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils), or At least some flowers unisexual. Fruit valves two. Fruit erect to spreading. Fruit segmentation segmented, or not segmented.

Albania, or Algeria, or Austria, or Belgium, or Canada, or Canary Islands, or Czech R, or Denmark, or Egypt, or Estonia, or Ethiopia, or Finland, or France, or Germany, or Greece, or Hungary, or Ireland, or Italy, or Japan, or Kenya, or Latvia, or Liberia, or Lithuania, or Mongolia, or Morocco, or Namibia, or Netherlands, or Norway, or Oman, or Poland, or Portugal, or Romania, or Russia, or Rwanda, or Saudi Arabia, or Slovakia, or Somalia, or Spain, or Sri Lanka, or Sudan, or Sweden, or Switzerland, or Tanzania, or Uganda, or United Kingdom, or United States, or Yemen, or Zimbabwe, or Congo, or Botswana.


Erysimum

Cheiranthus L.; Syrenia Andrz. ex Besser; Gynophorea Gilli; Syreniopsis H.P. Fuchs; Zederbauera H.P. Fuchs; Acachmena H. B. Fuchs.

Annual or biennial, or perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively malpighiaceous (2-branched, sessile, medifixed), or stellate (star-shaped) and sessile; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow, or purple (or violet), or orange; obovate, or oblong to elliptic, or orbicular, or spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free, or united. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or oblong; angustiseptate (flattened at a right angle to septum), or latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore at least 1.5 mm long, usually much longer, or absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50, or 51 to 100; Seeds uniseriate (in one row), or biseriate (in two rows); wing present, or absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina, or Ecuador, or Estonia, or Latvia, or Lithuania, or Belarus, or Kyrgyzstan, or Moldova, or Afghanistan, or Armenia, or Azerbaijan, or Bhutan, or China, or Cyprus, or Georgia, or India, or Iran, or Iraq, or Israel, or Japan, or Jordan, or Kazakhstan, or Korea, or Kyrgyzstan, or Lebanon, or Mongolia, or Nepal, or Pakistan, or Russia, or Saudi Arabia, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or Egypt, or Ukraine, or Czech R, or Slovakia, or Slovenia, or Costa Rica, or Guatemala, or Mexico, or Canada, or United States, or Greenland, or Australia, or Algeria, or Morocco, or Tunisia, or Canary Islands, or Cape Verde, or Madeira, or New Zealand, or Ireland, or United Kingdom, or Crete, or Greece, or Portugal, or Spain, or Albania, or Austria, or Belgium, or Balearic Islds, or France, or Germany, or Switzerland, or Netherlands, or Hungary, or Italy, or Yugoslavia, or Romania, or Sardinia, or Sicily, or Corsica, or Bulgaria, or Poland, or Croatia, or Macedonia, or Denmark, or Finland, or Iceland, or Norway, or Sweden.


Euclidium

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals subequalling sepals; white; spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovoid or obovoid; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick; indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages a beaklike style or distal segment. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Belarus, or Kyrgyzstan, or Moldova, or Afghanistan, or Armenia, or Azerbaijan, or China, or Georgia, or India, or Iran, or Iraq, or Jordan, or Kazakhstan, or Kyrgyzstan, or Lebanon, or Pakistan, or Russia, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or Uzbekistan, or Ukraine, or Czech R, or Slovakia, or Slovenia, or United States, or Austria, or Bulgaria, or Hungary, or Yugoslavia, or Romania.


Eudema

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves absent; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals; white; oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free, or united. Stamens six, or four; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); oblong, or ovate or obovate, or pyriform or obpyriform (pear-shaped); angustiseptate (flattened at a right angle to septum), or latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit two, or three to five, or six to ten, or eleven to 20, or 21 to 50, or 51 to 100; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina, or Bolivia, or Chile, or Ecuador, or Peru.


Eurycarpus

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched; multicellular glands absent; not a weed. Stem leaves absent; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; pink (or lavender), or purple (or violet); obovate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); lanceolate or oblanceolate, or oblong, or elliptic, or ovate or obovate; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds biseriate (in two rows); wing absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

China, or India, or Kazakhstan, or Pakistan.


Eutrema

Neomartinella Pilger; Platycraspedum O. E. Schulz; Taphrospermum C. A. Mey.; Thellungiella O. E. Schulz; Wasabia Matsum.

Annual or biennial, or perennial; herbs; terrestrial, or aquatic; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present, or absent; base petiolate, or sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or palmately lobed. Inflorescences racemes that are bracteate throughout or at least in lower half, or racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or absent; white, or pink (or lavender); obovate, or oblong to elliptic, or obcordate; margin and apex entire. Sepals ascending, or erect but not connivent, or spreading; free. Stamens six; lower part of filaments and petal claws without anything, or filaments with a wing, tooth, or appendage; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or lanceolate or oblanceolate, or oblong, or ovate or obovate, or cordate or obcordate; angustiseptate (flattened at a right angle to septum), or latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none, or wings. Ovules/seeds per ovary/fruit two, or three to five, or six to ten, or eleven to 20, or 21 to 50, or 51 to 100; Seeds uniseriate (in one row), or biseriate (in two rows); wing absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate, or whorled. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Kyrgyzstan, or Bhutan, or China, or India, or Japan, or Kazakhstan, or Korea, or Mongolia, or Nepal, or Russia, or Tajikistan, or Turkmenistan, or Turkey, or Uzbekistan, or Greenland, or United States, or Canada, or Taiwan.


Exhalimolobos

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple, or a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50, or 51 to 100, or more than 100; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Armenia, or Bolivia, or Colombia, or Ecuador, or Mexico, or Peru, or Uruguay, or Venezuela.


Farsetia

Annual or biennial, or perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively malpighiaceous (2-branched, sessile, medifixed); multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow, or purple (or violet); obovate, or oblong to elliptic, or orbicular, or spatulate to oblanceolate, or linear; margin and apex entire. Sepals erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or oblong, or elliptic, or ovate or obovate; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes decurrent (connivent); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row), or biseriate (in two rows); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or Bahrain, or Iran, or Iraq, or Israel, or Jordan, or Kuwait, or Oman, or Pakistan, or Qatar, or Saudi Arabia, or Syria, or United Arab Emir, or Yemen, or India, or Ethiopia, or Kenya, or Somalia, or Morocco, or Mauritania, or Algeria, or Tunisia, or Libya, or Chad, or Egypt, or Socotra, or Djibouti, or Mali, or Niger, or Sudan, or West Sahara, or Tanzania, or Uganda, or Bourkina Fasso, or Cameron, or Nigeria.


Fezia

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages wings. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation segmented.

Morocco.


Fibigia

Asterotricha V.V. Botsch., Irania Hadac & Chrtek; Pterygostemon V.V. Botsch.

Perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or stellate (star-shaped) and sessile; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow, or purple (or violet); spatulate to oblanceolate; margin and apex entire. Sepals erect and connivent; free. Stamens six; filaments with a wing, tooth, or appendage; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); oblong, or ovate or obovate, or orbicular; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds biseriate (in two rows); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Estonia, or Afghanistan, or Armenia, or Azerbaijan, or Cyprus, or Georgia, or Iran, or Iraq, or Israel, or Jordan, or Kazakhstan, or Lebanon, or Syria, or Turkmenistan, or Turkey, or Egypt, or Albania, or Yugoslavia, or Greece, or Bulgaria, or Crete, or Italy, or Ukraine, or Spain, or Austria, or France, or Bosnia & Herz, or Croatia, or Macedonia.


Foleyola

Perennial; shrubs or subshrubs; terrestrial; thorns and spines present; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; purple (or violet); oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or lanceolate or oblanceolate; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages none. Ovules/seeds per ovary/fruit three to five; Seeds uniseriate (in one row); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation segmented.

Algeria, or Morocco.


Fortuynia

Perennial; shrubs or subshrubs; terrestrial; thorns and spines present, or absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; purple (or violet); spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); orbicular, or cordate or obcordate; latiseptate (flattened parallel to septum); fruit (or segments) wall woody or corky and thick, or neither woody or papery (normal); indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages wings. Ovules/seeds per ovary/fruit two, or three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation segmented.

Afghanistan, or Iran, or Pakistan.


Fourraea

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand. Inflorescences racemes that are bracteate throughout or at least in lower half. Petals distinctly longer than sepals; white, or pink (or lavender); spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50, or 51 to 100; Seeds uniseriate (in one row); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Austria, or Belgium, or Corsica, or Czech R, or Slovakia, or France, or Germany, or Switzerland, or Spain, or Italy, or Yugoslavia, or Czech R, or Slovakia, or Slovenia.


Galitzkya

Perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) stellate (star-shaped) and sessile; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow; obovate; margin and apex bifid, 3-lobed, pinnately lobed, or fimbriate. Free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic, or ovate or obovate, or orbicular; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds biseriate (in two rows); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

China, or Kazakhstan, or Mongolia, or Russia.


Geococcus

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves absent; divisions and margins pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences solitary flowers borne on pedicels originating from basal rosette. Petals subequalling sepals; white; spatulate to oblanceolate; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row), or biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit reflexed (pendulous). Fruit segmentation not segmented.

Australia.


Glastaria

Texiera Jaub. & Spach.

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; obovate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); globose (spherical), or ovoid or obovoid; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick; indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Iraq, or Israel, or Jordan, or Lebanon, or Syria, or Turkey.


Glaucocarpum

Perennial; shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands present; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free, or united at least along lower third; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

United States.


Goldbachia

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands present, or absent; not a weed, or a weed. Stem leaves present; base sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet); spatulate to oblanceolate, or linear; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); lanceolate or oblanceolate, or ellipsoid; terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall woody or corky and thick; indehiscent (seeds remain inside whole fruit), or breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore at least 1.5 mm long, usually much longer, or absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed); fruit appendages none, or a beaklike style or distal segment. Ovules/seeds per ovary/fruit one, or two, or three to five; Seeds uniseriate (in one row), or aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Kyrgyzstan, or Afghanistan, or Armenia, or Azerbaijan, or China, or India, or Iran, or Iraq, or Kazakhstan, or Kyrgyzstan, or Mongolia, or Pakistan, or Russia, or Saudi Arabia, or Tajikistan, or Turkmenistan, or Turkey, or Uzbekistan.


Gorodkovia

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched; multicellular glands absent; not a weed. Stem leaves absent; divisions and margins 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); orbicular; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); lanceolate or oblanceolate, or oblong; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds biseriate (in two rows); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Russia.


Graellsia

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins palmately lobed. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate, or oblong to elliptic, or orbicular; margin and apex entire. Sepals spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); oblong; angustiseptate (flattened at a right angle to septum), or latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds uniseriate (in one row), or biseriate (in two rows), or aseriate (not in rows: when fruit has one or a few seeds); wing present, or absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or Iran, or Iraq, or Pakistan, or Tajikistan, or Turkmenistan, or Turkey, or Morocco, or Algeria, or Tunisia.


Grammosperma

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals subequalling sepals; white, or yellow; spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; petal claw and/or filament with hairs or papillae; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic, or orbicular; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina, or Chile.


Guiraoa

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; obovate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); globose (spherical), or ovoid or obovoid; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick, or neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages wings. Ovules/seeds per ovary/fruit three to five; Seeds uniseriate (in one row); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation segmented.

Spain.


Halimolobos

Poliophyton O. E. Schulz.

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or oblong, or ovoid or obovoid; angustiseptate (flattened at a right angle to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50, or 51 to 100, or more than 100; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Mexico, or United States, or Uruguay.


Harmsiodoxa

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); obovate, or orbicular; margin and apex entire. Sepals spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or fusiform; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row), or biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Australia.


Heldreichia

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); obovate; margin and apex entire. Sepals spreading; free. Stamens six; filaments with a wing, tooth, or appendage; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovate or obovate, or orbicular; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Lebanon, or Turkey.


Heliophila

Brachycarpaea DC.; Cycloptychis E. Mey. ex Sond.; Schlechteria Bolus ex Schltr.; Silicularia Compton; Thlaspeocarpa C. A. Sm.

Annual or biennial, or perennial; herbs, or shrubs or subshrubs, or lianas; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow, or pink (or lavender), or purple (or violet), or blue; obovate, or oblong to elliptic, or orbicular, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent, or spreading; free. Stamens six; lower part of filaments and petal claws without anything, or filaments with a wing, tooth, or appendage, or petal claw and/or filament with hairs or papillae; filaments of median stamens free; stamens included within calyx or partially exserted, or exserted well above sepals and petals. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or lanceolate or oblanceolate, or oblong, or elliptic, or ovate or obovate, or orbicular, or didymous (spectacle shaped); angustiseptate (flattened at a right angle to septum), or latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick, or neither woody or papery (normal); dehiscent (readily releasing seeds), or indehiscent (seeds remain inside whole fruit), or breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none, or a beaklike style or distal segment. Ovules/seeds per ovary/fruit one, or two, or three to five, or six to ten, or eleven to 20, or 21 to 50, or 51 to 100; Seeds uniseriate (in one row), or biseriate (in two rows), or aseriate (not in rows: when fruit has one or a few seeds); wing present, or absent; cotyledon diplecolobal, or spiral; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate, or opposite. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Australia, or South Africa, or Namibia, or Lesotho, or Swaziland.


Hemicrambe

Fabrisinapis Townsend; Nesocrambe A.G. Mill.

Perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow; obovate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or oblong; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds), or breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit two, or three to five, or six to ten; Seeds uniseriate (in one row); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation segmented.

Morocco, or Socotra.


Hemilophia

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple, or a mixture of simple and branched, or exclusively malpighiaceous (2-branched, sessile, medifixed); multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are bracteate throughout or at least in lower half. Petals distinctly longer than sepals; white, or yellow, or purple (or violet); obovate, or obcordate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything, or filaments with a wing, tooth, or appendage; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); oblong, or ovoid or obovoid; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed); fruit appendages crests. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

China.


Henophyton

Perennial; shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; purple (or violet); obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free, or united at least along lower third; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing present; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Algeria, or Libya, or Morocco, or Tunisia.


Hesperidanthus

Caulostramina Rollins; Glaucocarpum Rollins.

Perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands present, or absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow, or pink (or lavender), or purple (or violet); obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free, or united at least along lower third; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall thinly papery, or neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes decurrent (connivent); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50, or 51 to 100; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Mexico, or United States.


Hesperis

Diplopilosa F. Dvorák; Micrantha F. Dvorák.

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched; multicellular glands present; not a weed, or a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow, or purple (or violet), or green, or brown; obovate, or oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent, or erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Argentina, or Chile, or Estonia, or Latvia, or Lithuania, or Belarus, or Moldova, or Kyrgyzstan, or Armenia, or Azerbaijan, or China, or Georgia, or Iran, or Iraq, or Israel, or Japan, or Jordan, or Kazakhstan, or Kyrgyzstan, or Lebanon, or Mongolia, or Russia, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or Uzbekistan, or Canada, or United States, or Morocco, or Albania, or Yugoslavia, or Bulgaria, or Greece, or Ukraine, or Portugal, or Spain, or France, or Italy, or Sardinia, or Sicily, or Corsica, or Czech R, or Slovakia, or Hungary, or Romania, or Austria, or Poland, or Belgium, or United Kingdom, or Denmark, or Germany, or Switzerland, or Netherlands, or Finland, or Ireland, or Iceland, or Norway, or Sweden, or Czech R, or Slovakia, or Slovenia, or Mexico, or Canada, or United States, or Morocco, or New Zealand, or Ireland, or United Kingdom.


Hirschfeldia

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; obovate, or spatulate to oblanceolate; margin and apex entire. Sepals spreading, or reflexed; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens exserted well above sepals and petals. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall woody or corky and thick, or neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils), or At least some flowers unisexual. Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation segmented.

Albania, or Algeria, or Armenia, or Australia, or Austria, or Azerbaijan, or Belgium, or Canada, or Canary Islands, or Chile, or Cyprus, or Denmark, or Estonia, or France, or Georgia, or Germany, or Greece, or Hawaii, or Iran, or Iraq, or Ireland, or Israel, or Italy, or Japan, or Jordan, or Lebanon, or Latvia, or Libya, or Lithuania, or Morocco, or Netherlands, or New Zealand, or Norway, or Portugal, or Russia, or Saudi Arabia, or South Africa, or Spain, or Switzerland, or Syria, or Tunisia, or Ukraine, or Turkey, or Ukraine, or United Kingdom, or United States, or Vietnam, or Yemen.


Hollermayera

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; oblong to elliptic; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted, or exserted well above sepals and petals. Fruit a silicle (fruit up to 3 times longer than broad); pyriform or obpyriform (pear-shaped); latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Chile.


Hormathophylla

Perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines present, or absent; hairs (trichomes) stellate (star-shaped) and sessile; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow, or pink (or lavender); spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); orbicular; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit one, or two, or three to five, or six to ten; Seeds biseriate (in two rows), or aseriate (not in rows: when fruit has one or a few seeds); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Spain.


Hornungia

Hutchinsia R. Br.; Hutchinsiella O.E. Schulz; Hymenolobus Nutt. ex Torr. & A. Gray; Microcardamum O.E. Schulz; Pritzelago Kuntze.

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or stellate (star-shaped) and sessile, or at least some dendritic (tree-like); multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals; white; obovate, or oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or spreading; free. Stamens six, or four; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); lanceolate or oblanceolate, or ovate or obovate; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds biseriate (in two rows), or aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina, or Estonia, or Kyrgyzstan, or Afghanistan, or Armenia, or Azerbaijan, or China, or Cyprus, or Georgia, or India, or Iran, or Iraq, or Israel, or Jordan, or Kazakhstan, or Kyrgyzstan, or Japan, or Lebanon, or Mongolia, or Pakistan, or Russia, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or Uzbekistan, or Egypt, or Ukraine, or Slovakia, or Slovenia, or Mexico, or Canada, or United States, or Tasmania, or Australia, or Algeria, or Libya, or Morocco, or Tunisia, or New Zealand, or Canary Islands, or South Africa, or Bolivia, or Chile, or Ireland, or United Kingdom, or Crete, or Greece, or Portugal, or Spain, or Albania, or Austria, or Belgium, or Bosnia & Herz, or Bulgaria, or Corsica, or Croatia, or Denmark, or Finland, or France, or Germany, or Hungary, or Iceland, or Italy, or Macedonia, or Norway, or Poland, or Romania, or Sardinia, or Sicily, or Sweden, or Switzerland, or Yugoslavia.


Horwoodia

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; purple (or violet); oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything, or filaments with a wing, tooth, or appendage; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic, or orbicular; latiseptate (flattened parallel to septum); fruit (or segments) wall woody or corky and thick; indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages wings. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); cotyledon incumbent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit reflexed (pendulous). Fruit segmentation not segmented.

Iraq, or Kuwait, or Saudi Arabia.


Ianhedgea

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins palmately lobed, or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Kyrgyzstan, or Afghanistan, or China, or India, or Iran, or Pakistan, or Tajikistan, or Turkmenistan, or Uzbekistan.


Iberis

Annual or biennial, or perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet); oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovate or obovate, or orbicular, or cordate or obcordate; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages wings. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing present, or absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal), or zygomorphic (two petals and/or sepals much larger than other two); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Estonia, or Lithuania, or Moldova, or Cyprus, or Georgia, or Iran, or Iraq, or Israel, or Lebanon, or Syria, or Turkey, or Ukraine, or Czech R, or Slovakia, or Slovenia, or Canada, or United States, or Algeria, or Libya, or Morocco, or Tunisia, or New Zealand, or Canary Islands, or Ireland, or United Kingdom, or Crete, or Greece, or Portugal, or Spain, or Italy, or Sicily, or Romania, or Albania, or France, or Yugoslavia, or Switzerland, or Bulgaria, or Belgium, or Germany, or Netherlands, or Austria, or Bosnia & Herz, or Croatia, or Macedonia, or Poland, or Russia, or Balearic Islds.


Idahoa

Platyspermum Hook.

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves absent; divisions and margins entire or repand, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences solitary flowers borne on pedicels originating from basal rosette. Petals subequalling sepals; white; spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); orbicular; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds biseriate (in two rows); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

United States.


Iodanthus

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet); obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

United States.


Ionopsidium

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves absent; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate, or solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals; yellow, or pink (or lavender), or purple (or violet); obovate, or oblong to elliptic; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic, or ovate or obovate, or orbicular; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages wings. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Algeria, or Morocco, or Tunisia, or Greece, or Portugal, or Spain, or Ireland, or Italy, or Sicily.


Irenepharsus

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50; Seeds uniseriate (in one row), or biseriate (in two rows); wing present; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Australia.


Isatis

Goerkemia Yild.; Sameraria Desv.; Pachypterygium Bunge; Tauscheria Fisch. & C. A. Mey.

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed, or a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals; white, or yellow, or pink (or lavender); obovate, or oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or lanceolate or oblanceolate, or oblong, or ovate or obovate, or orbicular, or ellipsoid, or pyriform or obpyriform (pear-shaped), or cordate or obcordate, or navicular; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall woody or corky and thick, or neither woody or papery (normal); indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed); fruit appendages wings. Ovules/seeds per ovary/fruit one, or two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Estonia, or Latvia, or Lithuania, or Belarus, or Kyrgyzstan, or Moldova, or Afghanistan, or Armenia, or Azerbaijan, or China, or Georgia, or India, or Iran, or Iraq, or Israel, or Japan, or Jordan, or Kazakhstan, or Korea, or Kyrgyzstan, or Lebanon, or Mongolia, or Pakistan, or Russia, or Saudi Arabia, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or Uzbekistan, or Egypt, or Ukraine, or Czech R, or Slovakia, or Canada, or United States, or Algeria, or Libya, or Morocco, or Tunisia, or South Africa, or Canary Islands, or Madeira, or Argentina, or Peru, or Ireland, or United Kingdom, or Crete, or Greece, or Portugal, or Spain, or France, or Italy, or Bulgaria, or Hungary, or Yugoslavia, or Romania, or Albania, or Austria, or Belgium, or Bosnia & Herz, or Corsica, or Croatia, or Denmark, or Finland, or Germany, or Netherlands, or Macedonia, or Norway, or Poland, or Sardinia, or Sicily, or Sweden, or Switzerland.


Iskandera

Perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) stellate (star-shaped) and sessile, or at least some dendritic (tree-like); multicellular glands present; not a weed. Stem leaves absent; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; purple (or violet); obovate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Kyrgyzstan, or Tajikistan, or Uzbekistan.


Ivania

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore at least 1.5 mm long, usually much longer; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Chile.


Kernera

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate; margin and apex entire. Sepals spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); orbicular; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds biseriate (in two rows); wing present; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Slovenia, or Greece, or Spain, or Albania, or Austria, or Bosnia & Herz, or Bulgaria, or Corsica, or France, or Germany, or Switzerland, or Italy, or Croatia, or Macedonia, or Yugoslavia, or Poland, or Romania.


Kremeriella

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; obovate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ellipsoid; quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall woody or corky and thick; breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed); fruit appendages surface verrucose. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation segmented.

Algeria, or Morocco.


Lachnocapsa

Perennial; shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand. Inflorescences racemes that are bracteate throughout or at least in lower half. Petals distinctly longer than sepals; yellow; obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovate or obovate, or orbicular; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Socotra.


Lachnoloma

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovoid or obovoid; terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall woody or corky and thick; indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages spines. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Kyrgyzstan, or China, or Kazakhstan, or Iran, or Tajikistan, or Turkmenistan, or Uzbekistan.


Ladakiella

Perennial; herbs; terrestrial; thorns and spines present; hairs (trichomes) at least some dendritic (tree-like), or stellate and stalked; multicellular glands absent; not a weed. Stem leaves absent; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals; white, or pink (or lavender); spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovate or obovate, or ovoid or obovoid; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

China, or India.


Leavenworthia

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves absent; divisions and margins 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate, or solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals; white, or yellow, or pink (or lavender); obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or oblong, or cordate or obcordate; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing present; cotyledon accumbent, or straight; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

United States.


Leiocarpaea

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate; margin and apex entire. Sepals spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); globose (spherical), or ovoid or obovoid; terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall woody or corky and thick; indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon spiral; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Chile, or Kazakhstan, or Mongolia, or Russia.


Leiospora

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple, or a mixture of simple and branched; multicellular glands absent; not a weed. Stem leaves absent; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate, or solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals; pink (or lavender), or purple (or violet); obovate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50; Seeds uniseriate (in one row), or biseriate (in two rows); wing present, or absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Kyrgyzstan, or China, or India, or Kazakhstan, or Mongolia, or Pakistan, or Russia, or Tajikistan, or Uzbekistan.


Lepidium

Cardaria Desv.; Coronopus Zinn; Hymenophysa C.A. Mey.; Lepicochlea Rojas; Neolepia W.A. Weber; Papuzilla Ridl.; Senebiera DC.; Sprengeria Greene; Stoganowia Kar. & Kir.; Stubendorffia Schrenk ex Fisch. & C. A. Mey.; Winklera Regel.

Annual or biennial, or perennial; herbs, or shrubs or subshrubs, or lianas; terrestrial; thorns and spines present, or absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals, or reduced and smaller than sepals, or absent; white, or yellow, or pink (or lavender), or orange; obovate, or oblong to elliptic, or orbicular, or spatulate to oblanceolate, or linear, or filiform; margin and apex entire. Sepals ascending, or erect but not connivent, or spreading; free. Stamens six, or four, or two; lower part of filaments and petal claws without anything, or petal claw and/or filament with hairs or papillae; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); oblong, or elliptic, or ovate or obovate, or orbicular, or didymous (spectacle shaped), or globose (spherical), or cordate or obcordate; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall woody or corky and thick, or thinly papery, or neither woody or papery (normal); dehiscent (readily releasing seeds), or indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none, or wings, or surface verrucose. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing present, or absent; cotyledon accumbent, or incumbent, or diplecolobal; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils), or At least some flowers unisexual. Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina, or Bolivia, or Chile, or Colombia, or Ecuador, or Brazil, or Paraguay, or Peru, or Uruguay, or Venezuela, or Estonia, or Latvia, or Lithuania, or Belarus, or Kyrgyzstan, or Moldova, or Afghanistan, or Armenia, or Azerbaijan, or Bhutan, or China, or Cyprus, or Georgia, or India, or Indonesia, or Iran, or Iraq, or Israel, or Japan, or Jordan, or Kazakhstan, or Korea, or Kuwait, or Lebanon, or Mongolia, or Nepal, or Oman, or Pakistan, or Papua NG, or Russia, or Saudi Arabia, or Sri Lanka, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or United Arab Emir, or Uzbekistan, or Vietnam, or Yemen, or Egypt, or Ukraine, or Czech R, or Slovakia, or Slovenia, or Lesser Antilles, or Belize, or Canada, or Costa Rica, or El Salvador, or Guatemala, or Honduras, or Mexico, or Nicaragua, or Panama, or United States, or Australia, or Tasmania, or Tunisia, or Morocco, or Algeria, or Libya, or Kenya, or Tanzania, or Uganda, or Eritrea, or Ethiopia, or Zambia, or Zimbabwe, or Mozambique, or Congo Democratic R, or Madagascar, or Trinidad & Tobago, or Haiti, or Dominican R, or Cape Verde, or Sudan, or Botswana, or New Caledonia, or Taiwan, or Canary Islands, or Madeira, or Mali, or Nigeria, or Mauritania, or New Zealand, or Taiwan, or Hawaii, or Namibia, or South Africa, or Lesotho, or Swaziland, or Botswana, or Ireland, or United Kingdom, or Other Pacific Islands, or Other Indian Islands, or Other Atlantic Islands, or Comoros, or Socotra, or Crete, or Greece, or Portugal, or Spain, or Bahamas, or Cuba, or Haiti, or Puerto Rico, or Jamaica, or Trinidad & Tobago, or French Guiana, or Surinam, or Sudan, or Mozambique, or Zimbabwe, or Rwanda, or Bourkina Fasso, or Niger, or Guyana, or Other Caribbean Islands, or Dominican R, or Angola, or Togo, or Ivory Coast, or Central African R, or Congo, or Albania, or Austria, or Belgium, or Bosnia & Herz, or Bulgaria, or Corsica, or Croatia, or Denmark, or Finland, or France, or Germany, or Hungary, or Iceland, or Italy, or Macedonia, or Norway, or Poland, or Romania, or Sardinia, or Sicily, or Sweden, or Switzerland, or Yugoslavia, or Netherlands, or Gabon, or Cameron, or Guinea.


Lepidostemon

Chrysobraya H. Hara.

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present, or absent; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are bracteate throughout or at least in lower half, or racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate, or solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals; white, or yellow, or pink (or lavender), or purple (or violet); obovate; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything, or filaments with a wing, tooth, or appendage; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or oblong; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Bhutan, or China, or India, or Nepal.


Leptaleum

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate, or solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals; white, or pink (or lavender); spatulate to oblanceolate, or linear; margin and apex entire. Sepals erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens united at least along lower third; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick, or neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50, or 51 to 100, or more than 100; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Kyrgyzstan, or Afghanistan, or Armenia, or Azerbaijan, or China, or Georgia, or Iran, or Iraq, or Israel, or Jordan, or Kazakhstan, or Kuwait, or Pakistan, or Saudi Arabia, or Syria, or Tajikistan, or Turkmenistan, or Uzbekistan, or Egypt.


Lignariella

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or palmately lobed. Inflorescences racemes that are bracteate throughout or at least in lower half. Petals distinctly longer than sepals; pink (or lavender), or purple (or violet); obovate, or orbicular; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); oblong, or ellipsoid; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore at least 1.5 mm long, usually much longer, or absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit two, or three to five, or six to ten; Seeds uniseriate (in one row), or aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Bhutan, or China, or India, or Nepal.


Lithodraba

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves absent; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals; white; obovate; margin and apex entire. Sepals ascending; free. Stamens four; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovate or obovate; angustiseptate (flattened at a right angle to septum), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina.


Litwinowia

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands present; not a weed, or a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); globose (spherical); terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick; indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages a beaklike style or distal segment. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Kyrgyzstan, or Afghanistan, or Armenia, or Azerbaijan, or China, or Iran, or Kazakhstan, or Pakistan, or India, or Tajikistan, or Turkmenistan, or Uzbekistan.


Lobularia

Koniga R. Br.

Annual or biennial, or perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively malpighiaceous (2-branched, sessile, medifixed); multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet); obovate, or oblong to elliptic, or orbicular; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted, or exserted well above sepals and petals. Fruit a silicle (fruit up to 3 times longer than broad); elliptic, or ovate or obovate, or orbicular; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit two, or three to five, or six to ten, or eleven to 20; Seeds uniseriate (in one row), or biseriate (in two rows); wing present, or absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina, or Bolivia, or Ecuador, or Peru, or Estonia, or Lithuania, or Belarus, or Armenia, or China, or Cyprus, or Georgia, or Iran, or Iraq, or Israel, or Jordan, or Japan, or Lebanon, or Saudi Arabia, or Turkey, or Egypt, or Ukraine, or Czech R, or Slovakia, or Guatemala, or El Salvador, or Honduras, or Canada, or Mexico, or United States, or Australia, or Tasmania, or Algeria, or Libya, or Morocco, or Tunisia, or Canary Islands, or Madeira, or Eritrea, or Mozambique, or New Caledonia, or New Zealand, or Cape Verde, or Hawaii, or South Africa, or Brazil, or Chile, or Ireland, or United Kingdom, or Other Pacific Islands, or Other Atlantic Islands, or Other Indian Islands, or Greece, or Portugal, or Spain, or Albania, or Balearic Islds, or Corsica, or France, or Italy, or Sardinia, or Sicily, or Austria, or Belgium, or Denmark, or Netherlands, or Hungary, or Yugoslavia, or Bosnia & Herz, or Croatia, or Macedonia, or Norway, or Poland, or Romania.


Lunaria

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet); obovate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic, or ovate or obovate, or orbicular; latiseptate (flattened parallel to septum); fruit (or segments) wall thinly papery, or neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore at least 1.5 mm long, usually much longer; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten; Seeds biseriate (in two rows); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate, or opposite. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit reflexed (pendulous). Fruit segmentation not segmented.

Estonia, or Latvia, or Lithuania, or Afghanistan, or Moldova, or Georgia, or Japan, or Pakistan, or Turkey, or Ukraine, or Czech R, or Slovakia, or Slovenia, or Mexico, or Canada, or United States, or New Zealand, or Madeira, or Ireland, or United Kingdom, or Crete, or Greece, or Portugal, or Spain, or Albania, or Austria, or Belgium, or Bosnia & Herz, or Bulgaria, or Denmark, or France, or Germany, or Switzerland, or Hungary, or Italy, or Yugoslavia, or Croatia, or Macedonia, or Poland, or Romania, or Sardinia, or Sweden, or Corsica, or Netherlands, or Norway.


Lyrocarpa

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow, or purple (or violet), or brown; obovate, or spatulate to oblanceolate, or linear; margin and apex entire. Sepals erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); cordate or obcordate, or panduriform; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Mexico, or United States.


Macropodium

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or a mixture of simple and branched; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; spatulate to oblanceolate, or linear; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens exserted well above sepals and petals. Fruit a silique (fruit more than 3 times longer than broad); linear, or oblong; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore at least 1.5 mm long, usually much longer; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

China, or Japan, or Kazakhstan, or Mongolia, or Russia.


Malcolmia

Strigosella Boiss.

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or exclusively malpighiaceous (2-branched, sessile, medifixed), or stellate (star-shaped) and sessile; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet); obovate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50, or 51 to 100; Seeds uniseriate (in one row); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Albania, or Algeria, or Bosnia & Herz, or Bulgaria, or Cyprus, or France, or Greece, or Ireland, or Israel, or Italy, or Jordan, or Lebanon, or Morocco, or Portugal, or Saudi Arabia, or Spain, or Syria, or Tunisia, or Turkey, or United Kingdom, or Yugoslavia.


Mancoa

Hartwegiella O.E. Schulz; Poliophyton O.E. Schulz.

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or stellate (star-shaped) and sessile, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are bracteate throughout or at least in lower half, or racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals; white; obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted, or exserted well above sepals and petals. Fruit a silicle (fruit up to 3 times longer than broad); oblong, or elliptic; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50, or 51 to 100; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina, or Bolivia, or Mexico, or Peru, or United States.


Maresia

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) stellate (star-shaped) and sessile, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet); obovate, or spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Azerbaijan, or Cyprus, or Iran, or Iraq, or Israel, or Jordan, or Kuwait, or Lebanon, or Saudi Arabia, or Syria, or Turkey, or Algeria, or Libya, or Morocco, or Tunisia, or Albania, or Bulgaria, or Corsica, or Crete, or Greece, or France, or Spain, or Italy, or Sicily.


Matthiola

Lonchophora Durieu.

Annual or biennial, or perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or stellate (star-shaped) and sessile, or at least some dendritic (tree-like); multicellular glands present, or absent; not a weed, or a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow, or pink (or lavender), or purple (or violet), or brown; obovate, or oblong to elliptic, or spatulate to oblanceolate, or linear; margin and apex entire. Sepals erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything, or petal claw and/or filament with hairs or papillae; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages none, or wings, or horns. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50, or 51 to 100; Seeds uniseriate (in one row); wing present, or absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Chile, or Peru.


Mathewsia

Perennial; shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent, or erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or lanceolate or oblanceolate, or oblong, or elliptic; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Estonia, or Latvia, or Lithuania, or Kyrgyzstan, or Moldova, or Afghanistan, or Armenia, or Azerbaijan, or China, or Cyprus, or Georgia, or India, or Iran, or Iraq, or Israel, or Jordan, or Kazakhstan, or Kuwait, or Lebanon, or Pakistan, or Russia, or Saudi Arabia, or Syria, or Syria, or Turkmenistan, or Turkey, or Uzbekistan, or Egypt, or Ukraine, or Czech R, or Slovakia, or Slovenia, or United States, or Australia, or Tunisia, or Algeria, or Libya, or Morocco, or Kenya, or Ethiopia, or Somalia, or Djibouti, or Eritrea, or Canary Islands, or Madeira, or Mauritania, or New Zealand, or South Africa, or Lesotho, or Namibia, or Bolivia, or Ecuador, or Ireland, or United Kingdom, or Other Atlantic Islands, or Other Indian Islands, or Crete, or Greece, or Portugal, or Spain, or Djibouti, or Albania, or Corsica, or France, or Italy, or Yugoslavia, or Sardinia, or Sicily, or Balearic Islds, or Netherlands, or Bulgaria, or Switzerland.


Megacarpaea

Perennial; herbs; terrestrial; thorns and spines present, or absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or auriculate, amplexicaul, or sagittate; divisions and margins pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect, or pinnately compound. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals; yellow, or pink (or lavender), or purple (or violet); obovate, or spatulate to oblanceolate; margin and apex entire, or bifid, 3-lobed, pinnately lobed, or fimbriate. Sepals ascending, or erect but not connivent; free. Stamens six, or more than six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); didymous (spectacle shaped); angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages wings. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils), or At least some flowers unisexual. Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Kyrgyzstan, or Myanmar, or China, or India, or Kazakhstan, or Mongolia, or Nepal, or Pakistan, or Russia, or Tajikistan, or Turkmenistan, or Uzbekistan.


Megadenia

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present, or absent; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are bracteate throughout or at least in lower half, or solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals; white; obovate, or spatulate to oblanceolate; margin and apex entire. Sepals spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); didymous (spectacle shaped); angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

China, or Russia.


Menkea

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow, or pink (or lavender), or purple (or violet); obovate, or oblong to elliptic; margin and apex entire. Sepals spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); oblong, or elliptic; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50, or 51 to 100, or more than 100; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Australia.


Menonvillea

Hexaptera Hook.; Decaptera Turcz.

Annual or biennial, or perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple, or a mixture of simple and branched; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow, or pink (or lavender); obovate, or spatulate to oblanceolate, or linear; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything, or petal claw and/or filament with hairs or papillae; filaments of median stamens free, or united at least along lower third; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); didymous (spectacle shaped); angustiseptate (flattened at a right angle to septum); fruit (or segments) wall woody or corky and thick, or neither woody or papery (normal); breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore at least 1.5 mm long, usually much longer, or absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none, or wings. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina, or Chile.


Microlepidium

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals subequalling sepals, or absent; yellow; obovate; margin and apex entire. Sepals spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); orbicular, or triangular, or cordate or obcordate; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages wings. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Australia.


Microstigma

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) at least some dendritic (tree-like); multicellular glands present, or absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow, or pink (or lavender); oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or oblong, or ovoid or obovoid, or ellipsoid; latiseptate (flattened parallel to septum), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall woody or corky and thick; dehiscent (readily releasing seeds), or indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds uniseriate (in one row); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit reflexed (pendulous). Fruit segmentation not segmented.

China, or Mongolia, or Russia.


Microthlaspi

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed, or a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); orbicular, or cordate or obcordate; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages wings. Ovules/seeds per ovary/fruit three to five; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Belarus, or Kyrgyzstan, or Latvia, or Lithuania, or Afghanistan, or Armenia, or Azerbaijan, or China, or Cyprus, or Georgia, or Iran, or Iraq, or Israel, or Jordan, or Kazakhstan, or Pakistan, or Russia, or Saudi Arabia, or Tajikistan, or Turkmenistan, or Turkey, or Uzbekistan, or Ukraine, or Czech R, or Slovakia, or Slovenia, or United States, or Canada, or Algeria, or Libya, or Morocco, or Tunisia, or Somalia, or Ireland, or United Kingdom, or Crete, or Greece, or Portugal, or Spain, or Albania, or Austria, or Belgium, or Bosnia & Herz, or Bulgaria, or Corsica, or Croatia, or Denmark, or France, or Germany, or Hungary, or Italy, or Macedonia, or Poland, or Romania, or Sardinia, or Sicily, or Sweden, or Switzerland, or Yugoslavia.


Morettia

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) stellate (star-shaped) and sessile, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are bracteate throughout or at least in lower half. Petals distinctly longer than sepals, or subequalling sepals; white, or yellow, or pink (or lavender), or purple (or violet); obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); oblong; terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Bahrain, or Israel, or Jordan, or Oman, or Saudi Arabia, or United Arab Emir, or Yemen, or Egypt, or Libya, or Algeria, or Morocco, or Tunisia, or Mauritania, or Mali, or Niger, or Chad, or Sudan, or Ethiopia, or Somalia.


Moricandia

Oudneya R. Br.

Annual or biennial, or perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines present, or absent; hairs (trichomes) absent; multicellular glands absent; not a weed, or a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet); obovate, or spatulate to oblanceolate; margin and apex entire. Sepals erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50, or 51 to 100, or more than 100; Seeds uniseriate (in one row), or biseriate (in two rows); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Iran, or Israel, or Japan, or Jordan, or Oman, or Pakistan, or Saudi Arabia, or Yemen, or Egypt, or United States, or Algeria, or Libya, or Morocco, or Tunisia, or Mali, or Mauritania, or Canary Islands, or Greece, or Portugal, or Spain, or Balearic Islds, or Corsica, or Italy, or Sicily.


Moriera

Perennial; shrubs or subshrubs; terrestrial; thorns and spines present; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); oblong to elliptic; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic, or orbicular; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages wings. Ovules/seeds per ovary/fruit two, or three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or Iran, or Turkmenistan.


Morisia

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves absent; divisions and margins pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals; yellow; obovate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); lanceolate or oblanceolate, or oblong; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick, or neither woody or papery (normal); breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds uniseriate (in one row), or aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit reflexed (pendulous). Fruit segmentation segmented.

Corsica, or Sardinia.


Mostacillastrum

Phlebiophragmus O.E. Schulz.

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); oblong to elliptic; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50, or 51 to 100; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina, or Bolivia.


Murbeckiella

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); obovate, or oblong to elliptic; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50, or 51 to 100; Seeds uniseriate (in one row); wing present; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Armenia, or Azerbaijan, or Georgia, or Iran, or Turkey, or Morocco, or Portugal, or Spain, or France, or Sweden, or Italy.


Muricaria

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); obovate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); globose (spherical); terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick; indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages surface verrucose, or spines. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation segmented.

Algeria, or Libya, or Morocco, or Tunisia.


Myagrum

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed, or a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); pyriform or obpyriform (pear-shaped); angustiseptate (flattened at a right angle to septum); fruit (or segments) wall woody or corky and thick; indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Estonia, or Belarus, or Morocco, or Armenia, or Azerbaijan, or Georgia, or India, or Iran, or Iraq, or Israel, or Japan, or Jordan, or Kazakhstan, or Lebanon, or Syria, or Turkey, or Ukraine, or Czech R, or Slovakia, or Slovenia, or United States, or Canada, or Australia, or Algeria, or United Kingdom, or Greece, or Spain, or Albania, or Bulgaria, or France, or Germany, or Hungary, or Italy, or Yugoslavia, or Romania, or Sardinia, or Austria, or Switzerland, or Norway, or Sicily, or Sweden.


Nasturtiopsis

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; obovate; margin and apex entire. Sepals spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50, or 51 to 100; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Israel, or Jordan, or Egypt, or Libya, or Algeria, or Morocco, or Tunisia.


Nasturtium

Perennial; herbs; terrestrial, or aquatic; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or trifoliolate, or pinnately compound. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50; Seeds uniseriate (in one row), or biseriate (in two rows); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina, or Bolivia, or Ecuador, or Peru, or Estonia, or Lithuania, or Belarus, or Kyrgyzstan, or Moldova, or Myanmar, or Afghanistan, or Armenia, or Azerbaijan, or China, or Bhutan, or Cyprus, or Georgia, or India, or Indonesia, or Iran, or Iraq, or Israel, or Japan, or Jordan, or Kazakhstan, or Lebanon, or Malaysia, or Nepal, or Pakistan, or Papua NG, or Philippines, or Sri Lanka, or Saudi Arabia, or Syria, or Tajikistan, or Thailand, or Turkmenistan, or Turkey, or Uzbekistan, or Vietnam, or Yemen, or Russia, or Egypt, or Ukraine, or Czech R, or Slovakia, or Slovenia, or Lesser Antilles, or Canada, or Costa Rica, or El Salvador, or Guatemala, or Honduras, or Mexico, or Nicaragua, or Panama, or United States, or Cuba, or Australia, or Tasmania, or Algeria, or Libya, or Morocco, or Tunisia, or Kenya, or Tanzania, or Uganda, or Ethiopia, or Sudan, or Somalia, or Eritrea, or Cameron, or Congo Democratic R, or Lesotho, or Zambia, or Zimbabwe, or Taiwan, or Haiti, or Cuba, or Colombia, or Brazil, or Chile, or Uruguay, or Madeira, or Canary Islands, or Sierra Leon, or New Caledonia, or New Zealand, or Cape Verde, or Hawaii, or South Africa, or Lesotho, or Venezuela, or Chile, or Brazil, or Colombia, or Uruguay, or Ireland, or United Kingdom, or Other Pacific Islands, or Other Indian Islands, or Other Atlantic Islands, or Crete, or Greece, or Portugal, or Spain, or Cuba, or Puerto Rico, or Jamaica, or Trinidad & Tobago, or French Guiana, or Surinam, or Zimbabwe, or Mozambique, or Rwanda, or Dominican R, or Guyana, or Other Caribbean Islands, or Albania, or Austria, or Belgium, or Bosnia & Herz, or Bulgaria, or Corsica, or Croatia, or Denmark, or Finland, or France, or Germany, or Netherlands, or Hungary, or Italy, or Macedonia, or Poland, or Romania, or Sardinia, or Sicily, or Sweden, or Switzerland, or Yugoslavia, or Angola, or Mali, or Botswana, or Madagascar, or Bourkina Fasso, or Niger.


Neomartinella

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present, or absent; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate, or obcordate; margin and apex entire. Sepals spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or oblong; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

China.


Neotchihatechewia

Tchihatchewia Boiss.

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; pink (or lavender), or purple (or violet); oblong to elliptic, or orbicular; margin and apex entire. Sepals erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); orbicular; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages wings. Ovules/seeds per ovary/fruit three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Turkey.


Neotorularia

Torularia (Coss.) O.E. Schulz.

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present, or absent; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow, or pink (or lavender); obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or oblong; terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Kyrgyzstan, or Afghanistan, or Armenia, or Azerbaijan, or China, or Cyprus, or Georgia, or Iran, or Iraq, or Israel, or Jordan, or Kazakhstan, or Kuwait, or Lebanon, or Mongolia, or Pakistan, or Russia, or Saudi Arabia, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or Uzbekistan, or Egypt, or Algeria, or Libya, or Morocco, or Tunisia, or Spain.


Nerisyrenia

Greggia A. Gray.

Perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or oblong, or ovate or obovate; angustiseptate (flattened at a right angle to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50, or 51 to 100; Seeds uniseriate (in one row), or biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Mexico, or United States.


Neslia

Vogelia Medik.

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed, or a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); globose (spherical); terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick; indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit two, or three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina, or Estonia, or Latvia, or Lithuania, or Belarus, or Kyrgyzstan, or Moldova, or Afghanistan, or Armenia, or Azerbaijan, or China, or Cyprus, or Georgia, or India, or Iran, or Iraq, or Israel, or Japan, or Jordan, or Kazakhstan, or Kuwait, or Lebanon, or Mongolia, or Pakistan, or Russia, or Saudi Arabia, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or Uzbekistan, or Egypt, or Ukraine, or Czech R, or Slovakia, or Slovenia, or Canada, or United States, or Australia, or Algeria, or Libya, or Morocco, or Tunisia, or New Zealand, or Ireland, or United Kingdom, or Cuba, or Crete, or Greece, or Portugal, or Spain, or Albania, or Austria, or Balearic Islds, or Bulgaria, or Bosnia & Herz, or Corsica, or Croatia, or Denmark, or France, or Germany, or Hungary, or Italy, or Yugoslavia, or Macedonia, or Poland, or Romania, or Sardinia, or Sicily, or Sweden.


Neuontobotrys

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Argentina, or Chile, or Peru.


Nevada

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate, or spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

United States.


Noccaea

Brossardia Boiss.; Coluteocarpus Boiss.; Microthlaspi F. K. Mey.

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet); obovate, or oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or oblong, or elliptic, or ovate or obovate, or orbicular, or ovoid or obovoid, or ellipsoid, or cordate or obcordate; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall thinly papery, or neither woody or papery (normal); dehiscent (readily releasing seeds), or indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none, or wings. Ovules/seeds per ovary/fruit one, or two, or three to five, or six to ten, or eleven to 20; Seeds uniseriate (in one row), or biseriate (in two rows), or aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Afghanistan, or Albania, or Algeria, or Argentina, or Armenia, or Austria, or Azerbaijan, or Belarus, or Belgium, or Bhutan, or Bosnia & Herz, or Bulgaria, or Canada, or Chile, or China, or Corsica, or Croatia, or Cyprus, or Czech R, or Denmark, or Finland, or France, or Georgia, or Germany, or Greece, or Hungary, or Iceland, or India, or Iran, or Iraq, or Ireland, or Israel, or Italy, or Japan, or Jordan, or Kazakhstan, or Kyrgyzstan, or Lebanon, or Libya, or Lithuania, or Macedonia, or Mexico, or Mongolia, or Morocco, or Nepal, or Netherlands, or Norway, or Pakistan, or Poland, or Portugal, or Romania, or Russia, or Sardinia, or Saudi Arabia, or Sicily, or Slovakia, or Slovenia, or Somalia, or Spain, or Sweden, or Switzerland, or Syria, or Tajikistan, or Tunisia, or Turkmenistan, or Turkey, or Ukraine, or United Kingdom, or United States, or Uzbekistan, or Yugoslavia, or Crete.


Notoceras

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively malpighiaceous (2-branched, sessile, medifixed); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals subequalling sepals; yellow; spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); oblong; quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages horns. Ovules/seeds per ovary/fruit six to ten; Seeds uniseriate (in one row); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or India, or Iran, or Iraq, or Israel, or Jordan, or Kuwait, or Oman, or Pakistan, or Saudi Arabia, or Syria, or United Arab Emir, or Yemen, or Egypt, or Libya, or Tunisia, or Algeria, or Morocco, or Spain, or Canary Islands, or Mauritania.


Notothlaspi

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present, or absent; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic, or fusiform; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages wings. Ovules/seeds per ovary/fruit 21 to 50, or 51 to 100; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

New Zealand.


Ochthodium

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate; divisions and margins pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; obovate; margin and apex entire. Sepals spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); globose (spherical), or ellipsoid; quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall woody or corky and thick; indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages surface verrucose. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Cyprus, or Iraq, or Jordan, or Lebanon, or Syria, or Turkey, or Egypt.


Octoceras

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) stellate (star-shaped) and sessile, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals; white; obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); globose (spherical); quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall woody or corky and thick; indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages spines. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or Iran, or Kazakhstan, or Pakistan, or Turkmenistan, or Uzbekistan.


Olimarabidopsis

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively malpighiaceous (2-branched, sessile, medifixed), or stellate (star-shaped) and sessile, or stellate and stalked; multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals; yellow; oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50, or 51 to 100; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Afghanistan, or Armenia, or Azerbaijan, or China, or India, or Iran, or Iraq, or Israel, or Jordan, or Kazakhstan, or Kyrgyzstan, or Lebanon, or Oman, or Pakistan, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or Uzbekistan, or Egypt, or Russia.


Onuris

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand. Inflorescences racemes that are bracteate throughout or at least in lower half. Petals distinctly longer than sepals; white; obovate, or orbicular; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovate or obovate; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina, or Chile.


Oreoloma

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) at least some dendritic (tree-like); multicellular glands present; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow, or purple (or violet); obovate; margin and apex entire. Sepals erect but not connivent, or erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens united at least along lower third; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or oblong; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick; dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

China, or Mongolia.


Oreophyton

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves absent; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals; white, or pink (or lavender); obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Ethiopia, or Uganda, or Kenya, or Tanzania, or Rwanda.


Ornithocarpa

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); obovate, or orbicular; margin and apex bifid, 3-lobed, pinnately lobed, or fimbriate. Sepals spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); oblong, or ovate or obovate; latiseptate (flattened parallel to septum); fruit (or segments) wall woody or corky and thick, or neither woody or papery (normal); dehiscent (readily releasing seeds), or indehiscent (seeds remain inside whole fruit); gynophore at least 1.5 mm long, usually much longer, or absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none, or a beaklike style or distal segment. Ovules/seeds per ovary/fruit two, or six to ten, or eleven to 20; Seeds uniseriate (in one row), or aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Mexico.


Orychophragmus

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet); obovate, or obcordate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free, or united at least along lower third; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50, or 51 to 100; Seeds uniseriate (in one row); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

China, or Japan, or Mongolia.


Otocarpus

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; obovate, or oblong to elliptic; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic; latiseptate (flattened parallel to septum); fruit (or segments) wall woody or corky and thick; breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages wings. Ovules/seeds per ovary/fruit two, or three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation segmented.

Algeria.


Pachycladon

Cheesmania O.E. Schulz; Ischnocarpus O.E. Schulz.

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present, or absent; base petiolate; divisions and margins pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; angustiseptate (flattened at a right angle to septum), or latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row), or biseriate (in two rows); wing present, or absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils), or At least some flowers unisexual. Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

New Zealand, or Tasmania.


Pachymitus

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Australia.


Pachyneurum

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively malpighiaceous (2-branched, sessile, medifixed); multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or oblong; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Mongolia, or Russia.


Pachyphragma

Gagria Král.

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens exserted well above sepals and petals. Fruit a silicle (fruit up to 3 times longer than broad); cordate or obcordate; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall woody or corky and thick, or neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages wings. Ovules/seeds per ovary/fruit three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Azerbaijan, or Georgia, or Turkey.


Parlatoria

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate, or palmately lobed. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or elliptic; terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall woody or corky and thick, or neither woody or papery (normal); indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation segmented, or not segmented.

Iran, or Iraq, or Turkey.


Parodiodoxa

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals subequalling sepals; white; obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic; quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina.


Parolinia

Perennial; shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) stellate (star-shaped) and sessile, or at least some dendritic (tree-like), or stellate and stalked; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); oblong to elliptic, or spatulate to oblanceolate, or linear; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); oblong, or elliptic; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages horns. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds uniseriate (in one row); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Canary Islands.


Parrya

Achoriphragma Soják; Neuroloma Andrz. ex DC.

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands present, or absent; not a weed. Stem leaves present, or absent; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet); obovate, or spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or lanceolate or oblanceolate; latiseptate (flattened parallel to septum), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Kyrgyzstan, or Canada, or United States, or Afghanistan, or Bhutan, or China, or India, or Kazakhstan, or Pakistan, or Russia, or Tajikistan, or Uzbekistan.


Paysonia

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple, or a mixture of simple and branched; multicellular glands absent; not a weed, or a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow, or pink (or lavender); obovate; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); oblong, or ovate or obovate, or globose (spherical); angustiseptate (flattened at a right angle to septum), or latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick, or neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds biseriate (in two rows); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

United States.


Pegaeophyton

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves absent; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals; white, or pink (or lavender); obovate, or orbicular, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); oblong, or elliptic, or ovate or obovate, or orbicular; angustiseptate (flattened at a right angle to septum), or latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit two, or three to five, or six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Myanmar, or Bhutan, or China, or India, or Nepal, or Panama.


Peltaria

Leptoplax O.E. Schulz.

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate, or orbicular; margin and apex entire. Sepals spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic, or ovate or obovate, or orbicular; latiseptate (flattened parallel to septum); fruit (or segments) wall thinly papery, or neither woody or papery (normal); indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Iran, or Iraq, or Lebanon, or Syria, or Turkmenistan, or Turkey, or Greece, or Albania, or Austria, or Hungary, or Romania, or Yugoslavia, or Bosnia & Herz, or Croatia, or Macedonia.


Peltariopsis

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); oblong, or elliptic; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten; Seeds biseriate (in two rows); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Armenia, or Azerbaijan, or Iran, or Turkey.


Pennellia

Heterothrix (B.L. Rob.) Rydb.; Lamprophragma O.E. Schulz.

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals; white, or pink (or lavender), or purple (or violet); oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50, or 51 to 100, or more than 100; Seeds uniseriate (in one row), or biseriate (in two rows); wing present, or absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Argentina, or Bolivia, or Colombia, or Costa Rica, or Guatemala, or Mexico, or United States.


Petrocallis

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins palmately lobed. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); obovate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic, or ovate or obovate; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Iran, or Slovakia, or Slovenia, or Spain, or Austria, or France, or Germany, or Switzerland, or Italy, or Yugoslavia, or Romania.


Petroravenia

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves absent; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals reduced and smaller than sepals; white; linear; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovate or obovate; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina.


Phaeonychium

Parryopsis Botsch.; Vvedenskyella Botsch.; Wakilia Gilli.

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple, or a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present, or absent; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are bracteate throughout or at least in lower half, or racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet); obovate; margin and apex entire. Sepals ascending; free, or united. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or lanceolate or oblanceolate, or oblong, or ovoid or obovoid; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or Bhutan, or China, or India, or Nepal, or Pakistan, or Tajikistan.


Phlebolobium

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are bracteate throughout or at least in lower half. Petals distinctly longer than sepals; white; obovate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Falkland Isl.


Phlegmatospermum

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively malpighiaceous (2-branched, sessile, medifixed), or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow; obovate, or orbicular; margin and apex entire. Sepals spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic, or ovate or obovate, or orbicular; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Australia.


Phoenicaulis

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) stellate (star-shaped) and sessile, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; purple (or violet); obovate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or lanceolate or oblanceolate; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore at least 1.5 mm long, usually much longer, or absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Estonia, or United States.


Phravenia

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or oblong; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Mexico, or United States.


Physaria

Lesquerella S. Watson.

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) stellate (star-shaped) and sessile; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow, or orange; obovate, or oblong to elliptic, or spatulate to oblanceolate, or linear; margin and apex entire. Sepals ascending, or erect but not connivent, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); didymous (spectacle shaped), or globose (spherical), or ovoid or obovoid; angustiseptate (flattened at a right angle to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall thinly papery, or neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore at least 1.5 mm long, usually much longer, or absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row), or biseriate (in two rows); wing present, or absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Argentina, or Bolivia, or Mexico, or United States, or Canada, or Greenland, or Russia, or Canada.


Physocardamum

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or stellate (star-shaped) and sessile; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; oblong to elliptic; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); globose (spherical); angustiseptate (flattened at a right angle to septum); fruit (or segments) wall thinly papery; dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Turkey.


Physophtychis

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; obovate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; filaments with a wing, tooth, or appendage; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); globose (spherical); terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall thinly papery; dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds biseriate (in two rows), or aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Iran, or Iraq, or Turkey, or Armenia, or Azerbaijan.


Physorrhynchus

Perennial; shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); obovate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovoid or obovoid, or pyriform or obpyriform (pear-shaped); terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick; breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages none. Ovules/seeds per ovary/fruit two, or three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation segmented.

Afghanistan, or Iran, or Pakistan.


Planodes

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate; divisions and margins pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Mexico, or United States.


Platycraspedum

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or palmately lobed. Inflorescences racemes that are bracteate throughout or at least in lower half. Petals distinctly longer than sepals; white; obovate, or oblong to elliptic; margin and apex entire. Sepals ascending; free. Stamens six; filaments with a wing, tooth, or appendage; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or oblong; terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none, or wings. Ovules/seeds per ovary/fruit three to five, or six to ten; Seeds uniseriate (in one row); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

China.


Polyctenium

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); obovate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or oblong, or elliptic, or ovate or obovate; angustiseptate (flattened at a right angle to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

United States.


Polypsecadium

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit 51 to 100, or more than 100; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Argentina, or Bolivia, or Chile, or Colombia, or Ecuador, or Peru.


Pringlea

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand. Inflorescences racemes that are bracteate throughout or at least in lower half. Petals reduced and smaller than sepals, or absent; pink (or lavender); spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens exserted well above sepals and petals. Fruit a silicle (fruit up to 3 times longer than broad); ovoid or obovoid; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds biseriate (in two rows); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Other Indian Islands.


Pseuderucaria

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; purple (or violet); obovate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50, or 51 to 100, or more than 100; Seeds uniseriate (in one row); wing present; cotyledon accumbent, or conduplicate; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Egypt, or Israel, or Jordan, or Algeria, or Libya, or Morocco, or Tunisia.


Pseudoarabidopsis

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) stellate (star-shaped) and sessile; multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals; white, or pink (or lavender); spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit 51 to 100; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Ukraine, or Afghanistan, or China, or Kazakhstan, or Russia, or Tajikistan.


Pseudocamelina

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet); obovate, or spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or ellipsoid; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Iran.


Pseudoclausia

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands present; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; pink (or lavender), or purple (or violet); obovate, or spatulate to oblanceolate, or linear; margin and apex entire. Sepals erect but not connivent, or erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Kyrgyzstan, or Afghanistan, or China, or Iran, or Kazakhstan, or Tajikistan, or Turkmenistan, or Uzbekistan.


Pseudodraba

Perennial; herbs; terrestrial; thorns and spines present; hairs (trichomes) a mixture of simple and branched; multicellular glands absent; not a weed. Stem leaves present; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; obovate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); lanceolate or oblanceolate, or elliptic; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or Pakistan.


Pseudofortuynia

Perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; purple (or violet); obovate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore at least 1.5 mm long, usually much longer; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Iran.


Pseudosempervivum

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic, or ovate or obovate; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit two, or three to five; Seeds biseriate (in two rows), or aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Armenia, or Turkey.


Pseudoturritis

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) stellate and stalked; multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are bracteate throughout or at least in lower half. Petals distinctly longer than sepals; white, or yellow; spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit 51 to 100; Seeds uniseriate (in one row); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Albania, or Algeria, or Austria, or Belgium, or Bulgaria, or Czech R, or France, or Germany, or Greece, or Hungary, or Israel, or Italy, or Lebanon, or Romania, or Slovakia, or Spain, or Switzerland, or Syria, or Turkey, or Ukraine, or United Kingdom, or Yugoslavia.


Pseudovesicaria

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins palmately lobed. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); obovate, or orbicular; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten; Seeds biseriate (in two rows), or aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Armenia, or Azerbaijan, or Georgia.


Psychine

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are bracteate throughout or at least in lower half. Petals distinctly longer than sepals; white; obovate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); cordate or obcordate; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages wings. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50; Seeds biseriate (in two rows); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation segmented.

Algeria, or Libya, or Morocco.


Pterygiosperma

Perennial; shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; oblong to elliptic, or spatulate to oblanceolate, or linear; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; petal claw and/or filament with hairs or papillae; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing present; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina.


Pugionium

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); linear; margin and apex entire. Sepals erect but not connivent; united. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); lanceolate or oblanceolate, or oblong; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall woody or corky and thick, or neither woody or papery (normal); indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages wings, or spines. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

China, or Mongolia, or Russia.


Pycnoplinthopsis

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves absent; divisions and margins dentate, serrate, incised, crenate, sinuate. Inflorescences solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals; white; obovate; margin and apex entire. Sepals ascending; united. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or oblong; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Bhutan, or China, or India, or Nepal.


Pycnoplinthus

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves absent; divisions and margins entire or repand. Inflorescences solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals; white, or pink (or lavender); obovate; margin and apex entire. Sepals ascending; united. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

China, or India, or Pakistan.


Quezeliantha

Annual or biennial; herbs; terrestrial; thorns and spines present; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; purple (or violet); obovate, or oblong to elliptic; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything, or filaments with a wing, tooth, or appendage; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovate or obovate; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages none. Ovules/seeds per ovary/fruit six to ten; Seeds uniseriate (in one row); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Chad.


Raffenaldia

Cossonia Durieu.

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves absent; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals; yellow, or purple (or violet); obovate, or spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); oblong; quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall woody or corky and thick; indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten; Seeds uniseriate (in one row); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation segmented.

Algeria, or Morocco.


Raphanorrhyncha

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate, or orbicular; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); fusiform; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50; Seeds biseriate (in two rows); wing absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Mexico.


Raphanus

Quidproquo Greuter & Burdet.

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow, or pink (or lavender), or purple (or violet); obovate, or orbicular; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or oblong, or ellipsoid; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick; breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none, or a beaklike style or distal segment. Ovules/seeds per ovary/fruit two, or three to five, or six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation segmented, or not segmented.

Argentina, or Bolivia, or Ecuador, or Peru, or Estonia, or Latvia, or Lithuania, or Belarus, or Kyrgyzstan, or Moldova, or Myanmar, or Egypt, or Ukraine, or Afghanistan, or Armenia, or Azerbaijan, or Bahrain, or Bangladesh, or Bhutan, or Cambodia, or China, or Cyprus, or Georgia, or India, or Iran, or Iraq, or Indonesia, or Israel, or Japan, or Jordan, or Kazakhstan, or Korea, or Kuwait, or Laos, or Lebanon, or Malaysia, or Mongolia, or Nepal, or Oman, or Pakistan, or Papua NG, or Philippines, or Qatar, or Russia, or Sri Lanka, or Saudi Arabia, or Syria, or Tajikistan, or Thailand, or Turkmenistan, or Turkey, or United Arab Emir, or Uzbekistan, or Vietnam, or Yemen, or Czech R, or Slovakia, or Slovenia, or Lesser Antilles, or Belize, or Canada, or Costa Rica, or El Salvador, or Guatemala, or Honduras, or Mexico, or Nicaragua, or Panama, or United States, or Cuba, or Jamaica, or Haiti, or Dominican R, or Bahamas, or Puerto Rico, or Australia, or Tasmania, or Algeria, or Libya, or Morocco, or Tunisia, or Kenya, or Tanzania, or Burundi, or Eritrea, or Sudan, or Uganda, or Zambia, or Zimbabwe, or Malawi, or Canary Islands, or Mozambique, or Mali, or Nigeria, or New Caledonia, or New Zealand, or Madeira, or Ethiopia, or Taiwan, or Hawaii, or South Africa, or Colombia, or United Kingdom, or Ireland, or Other Pacific Islands, or Other Indian Islands, or Other Atlantic Islands, or Socotra, or Crete, or Greece, or Portugal, or Spain, or Bahamas, or Cuba, or Haiti, or Puerto Rico, or Surinam, or French Guiana, or Sudan, or Djibouti, or Zimbabwe, or Rwanda, or Guyana, or Other Caribbean Islands, or Dominican R, or Angola, or Togo, or Botswana, or Central African R, or Chad, or Congo, or Gabon, or Gambia, or Ghana, or Guinea, or Benin, or Liberia, or Mauritania, or Mauritius, or Sierra Leon, or Singapore, or Guinea Bissau, or Madagascar, or Albania, or Austria, or Belgium, or Bosnia & Herz, or Bulgaria, or Corsica, or Croatia, or Denmark, or Finland, or France, or Germany, or Hungary, or Iceland, or Italy, or Macedonia, or Netherlands, or Norway, or Poland, or Romania, or Sardinia, or Sicily, or Balearic Islds, or Sweden, or Switzerland, or Yugoslavia, or Cameron.


Rapistrum

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; obovate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); oblong, or ellipsoid; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick; breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit two, or three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation segmented.

Argentina, or Bolivia, or Brazil, or Chile, or Estonia, or Lithuania, or Belarus, or Moldova, or Egypt, or Ukraine, or Armenia, or Azerbaijan, or Cyprus, or Georgia, or India, or Iran, or Iraq, or Israel, or Japan, or Jordan, or Lebanon, or Pakistan, or Russia, or Saudi Arabia, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or Uzbekistan, or Yemen, or Czech R, or Slovakia, or Slovenia, or Canada, or Mexico, or United States, or Australia, or Tasmania, or Algeria, or Libya, or Morocco, or Tunisia, or Zambia, or Zimbabwe, or Canary Islands, or Madeira, or New Zealand, or Ethiopia, or South Africa, or United Kingdom, or Ireland, or Other Indian Islands, or Other Atlantic Islands, or Crete, or Greece, or Portugal, or Spain, or Albania, or Austria, or Belgium, or Bosnia & Herz, or Bulgaria, or Corsica, or Croatia, or France, or Germany, or Italy, or Macedonia, or Yugoslavia, or Netherlands, or Poland, or Romania, or Sardinia, or Sicily, or Switzerland.


Redowskia

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); orbicular, or fusiform; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit six to ten; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Russia.


Rhammatophyllum

Koeiea Rech. F.; Mitophyllum O.E. Schulz; Prionotrichon Botsch. & Vved.

Perennial; shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or exclusively malpighiaceous (2-branched, sessile, medifixed); multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow, or purple (or violet); obovate, or oblong to elliptic; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore at least 1.5 mm long, usually much longer, or absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing present; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Kyrgyzstan, or Afghanistan, or Kazakhstan, or Mongolia, or Tajikistan, or Turkmenistan.


Rhizobotrya

Perennial; shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are bracteate throughout or at least in lower half. Petals distinctly longer than sepals; white; obovate, or oblong to elliptic; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); globose (spherical), or ovoid or obovoid; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten; Seeds biseriate (in two rows); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Italy.


Ricotia

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect, or trifoliolate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet); obovate, or obcordate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds uniseriate (in one row); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit reflexed (pendulous). Fruit segmentation not segmented.

Egypt, or Armenia, or Israel, or Jordan, or Lebanon, or Syria, or Turkey, or Crete, or Greece.


Rollinsia

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; spatulate to oblanceolate; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; petal claw and/or filament with hairs or papillae; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore at least 1.5 mm long, usually much longer, or absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50, or 51 to 100; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Mexico.


Romanschulzia

Annual or biennial, or perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed, or a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals subequalling sepals, or reduced and smaller than sepals, or absent; white, or pink (or lavender); spatulate to oblanceolate, or linear; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore at least 1.5 mm long, usually much longer, or absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50, or 51 to 100; Seeds uniseriate (in one row), or biseriate (in two rows); wing absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Mexico, or Costa Rica, or Guatemala, or Panama.


Rorippa

Ceriosperma (O.E. Schulz) Greuter & Burdet; Kardanoglyphos Schlecht.; Neobeckia Greene; Roripella (Maire) Greuter & Burdet; Sisymbriella Spach; Trochiscus O.E. Schulz.

Annual or biennial, or perennial; herbs; terrestrial, or aquatic; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are bracteate throughout or at least in lower half, or racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals, or reduced and smaller than sepals, or absent; white, or yellow, or pink (or lavender); obovate, or oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or oblong, or globose (spherical), or ovoid or obovoid; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50, or 51 to 100, or more than 100; Seeds uniseriate (in one row), or biseriate (in two rows); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two, or three to six. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Argentina, or Bolivia, or Brazil, or Chile, or Colombia, or Ecuador, or Paraguay, or Peru, or Uruguay, or Estonia, or Latvia, or Lithuania, or Belarus, or Kyrgyzstan, or Moldova, or Myanmar, or Egypt, or Ukraine, or Afghanistan, or Armenia, or Azerbaijan, or Bangladesh, or Bhutan, or Cambodia, or China, or Georgia, or India, or Indonesia, or Iran, or Iraq, or Israel, or Japan, or Jordan, or Kazakhstan, or Korea, or Laos, or Lebanon, or Malaysia, or Mongolia, or Nepal, or Pakistan, or Papua NG, or Philippines, or Russia, or Sri Lanka, or Syria, or Syria, or Thailand, or Turkmenistan, or Turkey, or United Arab Emir, or Uzbekistan, or Vietnam, or Yemen, or Czech R, or Slovakia, or Slovenia, or Lesser Antilles, or Greenland, or Costa Rica, or Guatemala, or Honduras, or Nicaragua, or Panama, or Mexico, or Canada, or United States, or Cuba, or Tasmania, or Australia, or Algeria, or Libya, or Morocco, or Kenya, or Tanzania, or Uganda, or Ethiopia, or Congo Democratic R, or Rwanda, or Zambia, or Taiwan, or Zimbabwe, or Malawi, or Mozambique, or Madagascar, or Burundi, or Namibia, or Sudan, or Senegal, or Nigeria, or Ivory Coast, or Singapore, or New Caledonia, or Lesotho, or Somalia, or Mauritania, or New Zealand, or Taiwan, or Hawaii, or South Africa, or Mali, or Lesotho, or Venezuela, or Ireland, or United Kingdom, or Other Pacific Islands, or Other Indian Islands, or Other Atlantic Islands, or Comoros, or Greece, or Portugal, or Spain, or Bahamas, or Cuba, or Haiti, or Puerto Rico, or Jamaica, or Trinidad & Tobago, or French Guiana, or Surinam, or Sudan, or Zambia, or Zimbabwe, or Rwanda, or Guyana, or Other Caribbean Islands, or Dominican R, or Albania, or Austria, or Belgium, or Bosnia & Herz, or Bulgaria, or Corsica, or Croatia, or Denmark, or Finland, or France, or Germany, or Netherlands, or Hungary, or Iceland, or Italy, or Macedonia, or Norway, or Poland, or Romania, or Sardinia, or Sicily, or Sweden, or Switzerland, or Yugoslavia, or Angola, or Ivory Coast, or Congo, or Gabon, or Mauritius, or Sierra Leon, or Cameron.


Rytidocarpus

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; purple (or violet); obovate, or orbicular; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); oblong, or elliptic; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages a beaklike style or distal segment. Ovules/seeds per ovary/fruit 21 to 50; Seeds biseriate (in two rows); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation segmented.

Morocco.


Sandbergia

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) stellate and stalked; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); At least some flowers unisexual. Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

United States.


Sarcodraba

Ateixa Ravenna; Grammosperma O. E. Schulz.

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or a mixture of simple and branched; multicellular glands absent; not a weed. Stem leaves present, or absent; base petiolate, or sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals; white, or yellow; obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; petal claw and/or filament with hairs or papillae; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or oblong, or elliptic, or ovate or obovate, or orbicular; angustiseptate (flattened at a right angle to septum), or latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina, or Chile.


Savignya

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet); obovate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds biseriate (in two rows); wing present; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Egypt, or Afghanistan, or Bahrain, or Iran, or Iraq, or Israel, or Jordan, or Kuwait, or Oman, or Pakistan, or Qatar, or Saudi Arabia, or United Arab Emir, or Algeria, or Libya, or Morocco, or Tunisia.


Scambopus

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; obovate, or orbicular; margin and apex entire. Sepals spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); fusiform; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50; Seeds uniseriate (in one row), or biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Australia.


Schimpera

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals subequalling sepals; yellow; spatulate to oblanceolate; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic, or orbicular; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick; indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed); fruit appendages a beaklike style or distal segment. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Egypt, or Iran, or Iraq, or Israel, or Jordan, or Kuwait, or Qatar, or Saudi Arabia, or Syria, or United Arab Emir.


Schivereckia

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate; margin and apex entire. Sepals ascending; free. Stamens six; filaments with a wing, tooth, or appendage; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); oblong, or ovate or obovate; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds biseriate (in two rows); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Moldova, or Ukraine, or Russia, or Turkey, or Romania, or Albania, or Yugoslavia.


Schizopetalon

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate; margin and apex bifid, 3-lobed, pinnately lobed, or fimbriate. Sepals erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50, or 51 to 100; Seeds uniseriate (in one row), or biseriate (in two rows); wing absent; cotyledon incumbent, or of other types; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Argentina, or Chile.


Schlechteria

Perennial; shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow; obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic, or ovate or obovate; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit two, or three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon diplecolobal; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit reflexed (pendulous). Fruit segmentation not segmented.

South Africa.


Schouwia

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; purple (or violet); obovate, or oblong to elliptic; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); orbicular, or didymous (spectacle shaped); angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages wings, or a beaklike style or distal segment. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation segmented.

Egypt, or India, or Jordan, or Saudi Arabia, or Yemen, or Libya, or Algeria, or Morocco, or Somalia, or Mali, or Mauritania, or Chad, or Niger, or Sudan, or Ethiopia, or Djibouti, or Bourkina Fasso.


Schrenkiella

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals subequalling sepals, or reduced and smaller than sepals, or absent; white; spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Armenia, or Azerbaijan, or Chile, or Iran, or Kazakhstan, or Turkmenistan, or Turkey, or Uzbekistan.


Scoliaxon

Perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); globose (spherical); terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten; Seeds biseriate (in two rows), or aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Mexico.


Selenia

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are bracteate throughout or at least in lower half, or solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals; yellow; obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); oblong, or globose (spherical); latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds biseriate (in two rows); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Mexico, or United States.


Shangrilaia

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals; white; spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovoid or obovoid; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

China.


Sibara

Machaeorphorus Schltd.; Pterygiosperma O. E. Schulz; Werdermannia O. E. Schulz.

Annual or biennial, or perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple, or a mixture of simple and branched; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals; white, or pink (or lavender), or purple (or violet); obovate, or oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything, or petal claw and/or filament with hairs or papillae; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50, or 51 to 100; Seeds uniseriate (in one row), or biseriate (in two rows); wing present, or absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Argentina, or Chile, or Mexico, or United States.


Sibaropsis

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; pink (or lavender); spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens united at least along lower third; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

United States.


Silicularia

Perennial; shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); obovate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic, or ovate or obovate, or orbicular; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon diplecolobal; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit reflexed (pendulous). Fruit segmentation not segmented.

South Africa.


Sinapidendron

Perennial; shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; oblong to elliptic; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50, or 51 to 100; Seeds uniseriate (in one row); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Madeira.


Sinapis

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow; obovate; margin and apex entire. Sepals spreading, or reflexed; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or oblong; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none, or a beaklike style or distal segment. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation segmented.

Argentina, or Bolivia, or Estonia, or Latvia, or Lithuania, or Belarus, or Kyrgyzstan, or Moldova, or Egypt, or Ukraine, or Afghanistan, or Armenia, or Azerbaijan, or China, or Cyprus, or India, or Iran, or Iraq, or Israel, or Japan, or Jordan, or Kazakhstan, or Korea, or Lebanon, or Mongolia, or Pakistan, or Russia, or Saudi Arabia, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or Uzbekistan, or Vietnam, or Czech R, or Slovakia, or Slovenia, or Lesser Antilles, or Canada, or Mexico, or United States, or Australia, or Tasmania, or Algeria, or Libya, or Morocco, or Tunisia, or Canary Islands, or New Zealand, or Madeira, or South Africa, or Colombia, or Brazil, or Ireland, or United Kingdom, or Other Atlantic Islands, or Crete, or Greece, or Portugal, or Spain, or Other Caribbean Islands, or Bahamas, or Cuba, or Haiti, or Puerto Rico, or Zimbabwe, or Togo, or Ivory Coast, or Albania, or Austria, or Belgium, or Bosnia & Herz, or Bulgaria, or Corsica, or Croatia, or Denmark, or Balearic Islds, or Finland, or France, or Germany, or Hungary, or Iceland, or Italy, or Macedonia, or Netherlands, or Norway, or Poland, or Romania, or Sardinia, or Sicily, or Sweden, or Switzerland, or Yugoslavia, or Gabon.


Sisymbrella

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or auriculate, amplexicaul, or sagittate; divisions and margins 1–3-pinnatisect. Inflorescences racemes that are bracteate throughout or at least in lower half, or racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; obovate, or spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50; Seeds biseriate (in two rows); wing present, or absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Algeria, or France, or Italy, or Morocco, or Portugal, or Spain.


Sisymbriopsis

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are bracteate throughout or at least in lower half, or racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); obovate, or spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing present, or absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Kyrgyzstan, or China, or Tajikistan.


Sisymbrium

Lycocarpus O.E. Schulz; Phryne Bubani; Schoenocrambe Greene.

Annual or biennial, or perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are bracteate throughout or at least in lower half, or racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals; white, or yellow, or pink (or lavender); obovate, or oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent, or spreading; free, or united. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or lanceolate or oblanceolate; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50, or 51 to 100, or more than 100; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Argentina, or Bolivia, or Chile, or Ecuador, or Peru, or Estonia, or Latvia, or Lithuania, or Belarus, or Kyrgyzstan, or Moldova, or Egypt, or Ukraine, or Afghanistan, or Armenia, or Azerbaijan, or Bahrain, or China, or Cyprus, or Georgia, or India, or Iran, or Iraq, or Israel, or Japan, or Jordan, or Kazakhstan, or Korea, or Kuwait, or Lebanon, or Mongolia, or Nepal, or Oman, or Pakistan, or Qatar, or Russia, or Saudi Arabia, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or United Arab Emir, or Uzbekistan, or Yemen, or Czech R, or Slovakia, or Slovenia, or Lesser Antilles, or Costa Rica, or Mexico, or Canada, or United States, or Australia, or Tasmania, or Algeria, or Libya, or Morocco, or Tunisia, or Kenya, or Tanzania, or Ethiopia, or Sudan, or Canary Islands, or Madeira, or Brazil, or Socotra, or Somalia, or Eritrea, or Zambia, or Zimbabwe, or Mauritania, or New Caledonia, or New Zealand, or Taiwan, or Hawaii, or South Africa, or Namibia, or Lesotho, or Brazil, or Ireland, or United Kingdom, or Other Pacific Islands, or Other Atlantic Islands, or Other Indian Islands, or Cuba, or Haiti, or Jamaica, or Djibouti, or Zambia, or Niger, or Dominican R, or Other Caribbean Islands, or Crete, or Greece, or Portugal, or Spain, or Belgium, or France, or Germany, or Switzerland, or Netherlands, or Sweden, or Albania, or Australia, or Bulgaria, or Hungary, or Italy, or Yugoslavia, or Poland, or Romania, or Corsica, or Sardinia, or Sicily, or Denmark, or Balearic Islds.


Smelowskia

Acroschizocarpus Gombocz; Chrysthemopsis Rech. f.; Gorodkovia Botsch. & Karav.; Hedinia Ostenf.; Hediniopsis Botsch. & Petrovsky; Melanidion Greene; Redowskia Cham. & Schltdl.; Sinosophiopsis Al-Shehbaz; Sophiopsis O. E. Schulz.

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present, or absent; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are bracteate throughout or at least in lower half, or racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow, or pink (or lavender); obovate, or oblong to elliptic, or orbicular, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or lanceolate or oblanceolate, or oblong, or elliptic, or ovate or obovate, or orbicular, or ovoid or obovoid, or ellipsoid, or pyriform or obpyriform (pear-shaped), or fusiform; angustiseptate (flattened at a right angle to septum), or latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row), or biseriate (in two rows); wing present, or absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Kyrgyzstan, or Afghanistan, or Bhutan, or China, or India, or Kazakhstan, or Mongolia, or Nepal, or Pakistan, or Russia, or Tajikistan, or Canada, or United States, or Uzbekistan.


Sobolewskia

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate, or palmately lobed. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); oblong, or ellipsoid; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick; indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Armenia, or Azerbaijan, or Georgia, or Turkey, or Ukraine.


Solms-laubachia

Desideria Pamp.; Ermaniopsis H. Hara; Eurycarpus Botsch.; Oreoblastus Suslova; Parryopsis Botsch.; Phaeonychium O.E. Schulz; Vvedenskyella Botsch.; Wakilia Gilli.

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple, or a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present, or absent; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid), or palmately lobed. Inflorescences racemes that are bracteate throughout or at least in lower half, or racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate, or solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet), or blue; obovate, or oblong to elliptic, or orbicular, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent, or erect and connivent; free, or united. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or lanceolate or oblanceolate, or oblong, or elliptic, or ovate or obovate, or ovoid or obovoid; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20, or 21 to 50, or 51 to 100; Seeds uniseriate (in one row), or biseriate (in two rows); wing absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or Bhutan, or Chile, or China, or India, or Kazakhstan.


Sphaerocardamum

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or absent; white; spatulate to oblanceolate; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted, or exserted well above sepals and petals. Fruit a silicle (fruit up to 3 times longer than broad); oblong, or globose (spherical); angustiseptate (flattened at a right angle to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row), or biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Mexico.


Spirorhynchus

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands present, or absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); spatulate to oblanceolate, or linear; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); ellipsoid; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick; indehiscent (seeds remain inside whole fruit); gynophore at least 1.5 mm long, usually much longer; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed); fruit appendages a beaklike style or distal segment. Ovules/seeds per ovary/fruit one, or two; Seeds aseriate (not in rows: when fruit has one or a few seeds); cotyledon incumbent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Afghanistan, or China, or Iran, or Kazakhstan, or Kyrgyzstan, or Pakistan, or Tajikistan, or Turkmenistan, or Uzbekistan.


Spryginia

Petiniotia J. Léonard; Sterigma DC.

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple, or a mixture of simple and branched; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; pink (or lavender), or purple (or violet); obovate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens united at least along lower third; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50, or 51 to 100, or more than 100; Seeds uniseriate (in one row); wing present; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Afghanistan, or Tajikistan, or Turkmenistan, or Uzbekistan.


Stanleya

Annual or biennial, or perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow; oblong to elliptic, or spatulate to oblanceolate, or linear; margin and apex entire. Sepals spreading, or reflexed; free. Stamens six; petal claw and/or filament with hairs or papillae; filaments of median stamens free; stamens exserted well above sepals and petals. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore at least 1.5 mm long, usually much longer; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50, or 51 to 100, or more than 100; Seeds uniseriate (in one row); wing absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

United States.


Stenopetalum

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or exclusively malpighiaceous (2-branched, sessile, medifixed); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or orange, or green, or brown; linear, or filiform; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free, or united at least along lower third; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or oblong, or ovate or obovate, or orbicular; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick, or neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds biseriate (in two rows); wing absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Australia, or Tasmania.


Sterigmostemum

Oreoloma Botsch.

Annual or biennial, or perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands present; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow, or purple (or violet), or orange; obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent, or erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free, or united at least along lower third; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or oblong; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall woody or corky and thick; breaks up into closed, 1- or few-seeded segments (schizocarp, loment, or segmented); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none, or wings, or spines. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Kyrgyzstan, or Armenia, or Azerbaijan, or China, or Georgia, or Iran, or Iraq, or Kazakhstan, or Lebanon, or Mongolia, or Pakistan, or Russia, or Syria, or Turkmenistan, or Turkey, or Yemen.


Stevenia

Berteroella O. E. Schulz.

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) stellate (star-shaped) and sessile; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet); obovate, or oblong to elliptic, or orbicular; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or oblong, or elliptic; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit two, or three to five, or six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing present, or absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

China, or Japan, or Kazakhstan, or Korea, or Mongolia, or Russia.


Straussiella

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) stellate (star-shaped) and sessile; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow, or purple (or violet); obovate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; filaments with a wing, tooth, or appendage; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovoid or obovoid; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten; Seeds biseriate (in two rows); wing present; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Iran.


Streptanthella

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens exserted well above sepals and petals. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing present; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

United States.


Streptanthus

Agianthus Greene; Cartiera Greene; Caulanthus S. Watson; Disaccanthus Greene; Euklisia Rydb. ex Small; Icianthus Greene; Mesoreanthus Greene; Microsemia Greene; Mitophyllum Greene; Pleiocardia Greene; Sibaropsis S. Boyd & T. S. Ross; Streptanthella Rydb.

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals; white, or yellow, or pink (or lavender), or purple (or violet); obovate, or oblong to elliptic, or spatulate to oblanceolate, or linear; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free, or united at least along lower third; stamens included within calyx or partially exserted, or exserted well above sepals and petals. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore at least 1.5 mm long, usually much longer, or absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50, or 51 to 100, or more than 100; Seeds uniseriate (in one row); wing present, or absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal), or zygomorphic (two petals and/or sepals much larger than other two); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Mexico, or United States.


Streptoloma

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively malpighiaceous (2-branched, sessile, medifixed); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); obovate, or oblong to elliptic; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or Iran, or Jordan, or Kyrgyzstan, or Tajikistan, or Turkmenistan, or Uzbekistan.


Strigosella

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed, or a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet); oblong to elliptic, or obcordate; margin and apex entire. Sepals erect but not connivent, or erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free, or united at least along lower third; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50, or 51 to 100; Seeds uniseriate (in one row); wing present, or absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Argentina, or Estonia, or Belarus, or Kazakhstan, or Kyrgyzstan, or Afghanistan, or Armenia, or Azerbaijan, or China, or Cyprus, or Georgia, or India, or Iran, or Iraq, or Israel, or Jordan, or Kazakhstan, or Kuwait, or Mongolia, or Oman, or Pakistan, or Lebanon, or Qatar, or Russia, or Saudi Arabia, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or Uzbekistan, or United Arab Emir, or Egypt, or Ukraine, or Czech R, or Slovakia, or United States, or Canada, or Algeria, or Libya, or Morocco, or Tunisia, or New Zealand, or Canary Islands, or Madeira, or Ireland, or United Kingdom, or Crete, or Greece, or Portugal, or Spain, or France, or Italy, or Balearic Islds, or Corsica, or Sardinia, or Sicily, or Hungary, or Albania, or Yugoslavia.


Subularia

Annual or biennial; herbs; aquatic; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves absent; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals, or absent; white; oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovoid or obovoid, or ellipsoid; angustiseptate (flattened at a right angle to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Estonia, or Latvia, or Lithuania, or Belarus, or Ukraine, or Mongolia, or Russia, or Canada, or United States, or Kenya, or Tanzania, or Uganda, or Ethiopia, or Congo Democratic R, or Ireland, or United Kingdom, or Spain, or Rwanda, or Belgium, or Bulgaria, or Denmark, or Finland, or France, or Germany, or Netherlands, or Iceland, or Norway, or Sweden.


Succowia

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; obovate, or oblong to elliptic; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovate or obovate, or orbicular; angustiseptate (flattened at a right angle to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages spines. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Algeria, or Morocco, or Tunisia, or Spain, or Corsica, or Sardinia, or Sicily, or Italy, or Canary Islands, or Balearic Islds.


Synstemon

Synstemonanthus Botsch.

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals subequalling sepals; pink (or lavender); obovate; margin and apex entire. Sepals ascending; free. Stamens six; petal claw and/or filament with hairs or papillae; filaments of median stamens united at least along lower third; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing present; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

China.


Synthlipsis

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); obovate; margin and apex entire. Sepals spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); oblong, or elliptic; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing present, or absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Mexico, or United States.


Tchihatechewia

Neotchihatchewia.

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are bracteate throughout or at least in lower half, or racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; pink (or lavender), or purple (or violet); oblong to elliptic, or orbicular; margin and apex entire. Sepals erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); orbicular; angustiseptate (flattened at a right angle to septum), or latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages wings. Ovules/seeds per ovary/fruit three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Turkey.


Teesdalia

Teesdaliopsis (Willk.) Rothm.

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed, or a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals; white; obovate, or oblong to elliptic; margin and apex entire. Sepals ascending, or spreading; free. Stamens six, or four; filaments with a wing, tooth, or appendage; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); orbicular, or cordate or obcordate; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages wings. Ovules/seeds per ovary/fruit two, or three to five; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers zygomorphic (two petals and/or sepals much larger than other two); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Estonia, or Lithuania, or Belarus, or Ukraine, or Cyprus, or Lebanon, or Turkey, or Slovakia, or Canada, or United States, or Tasmania, or Algeria, or Morocco, or Tunisia, or Madeira, or Ireland, or United Kingdom, or Crete, or Greece, or Portugal, or Spain, or Austria, or Belgium, or Bulgaria, or Corsica, or Denmark, or France, or Germany, or Switzerland, or Netherlands, or Hungary, or Italy, or Yugoslavia, or Norway, or Poland, or Romania, or Russia, or Sicily, or Sweden, or Sardinia.


Tetracme

Tetracmidion Korsh.

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) stellate (star-shaped) and sessile, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals; white; obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or oblong, or ovoid or obovoid; terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall woody or corky and thick, or neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim, or complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages horns. Ovules/seeds per ovary/fruit two, or three to five, or six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Kyrgyzstan, or Afghanistan, or China, or India, or Iran, or Kazakhstan, or Mongolia, or Pakistan, or Tajikistan, or Turkmenistan, or Uzbekistan, or Russia.


Thellungiella

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or absent; white; obovate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50, or 51 to 100; Seeds uniseriate (in one row), or biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Armenia, or Azerbaijan, or China, or Iran, or Kazakhstan, or Kyrgyzstan, or Mongolia, or Russia, or Turkmenistan, or Turkey, or Uzbekistan, or Canada, or United States.


Thelypodiopsis

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or yellow, or pink (or lavender), or purple (or violet); obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted, or exserted well above sepals and petals. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore at least 1.5 mm long, usually much longer, or absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50, or 51 to 100; Seeds uniseriate (in one row); wing absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Mexico, or United States.


Thelypodium

Pachypodium Nutt. ex Torr. & A. Gray; Pleurophragma Rydb.; Stanleyella Rydb.

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet); obovate, or spatulate to oblanceolate, or linear; margin and apex entire. Sepals ascending, or erect but not connivent, or spreading, or reflexed; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free, or united at least along lower third; stamens included within calyx or partially exserted, or exserted well above sepals and petals. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore at least 1.5 mm long, usually much longer, or absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit eleven to 20, or 21 to 50, or 51 to 100; Seeds uniseriate (in one row); wing absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Canada, or Mexico, or United States.


Thlaspeocarpa

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins entire or repand, or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; obovate, or oblong to elliptic; margin and apex entire. Sepals ascending, or spreading; free. Stamens six; filaments with a wing, tooth, or appendage; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic, or ovate or obovate, or orbicular; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit one, or two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon diplecolobal; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate, or opposite. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit reflexed (pendulous). Fruit segmentation not segmented.

South Africa.


Thlaspi

Gagria Král; Pachyphragma (DC.) Rchb.

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed, or a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals; white; obovate, or oblong to elliptic; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); oblong, or ovate or obovate, or cordate or obcordate; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none, or wings, or horns. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Armenia, or Azerbaijan, or Turkey, or Argentina, or Chile, or Estonia, or Latvia, or Lithuania, or Belarus, or Kyrgyzstan, or Moldova, or Ukraine, or Afghanistan, or Bhutan, or China, or Georgia, or Cyprus, or India, or Iran, or Israel, or Japan, or Kazakhstan, or Korea, or Lebanon, or Mongolia, or Nepal, or Pakistan, or Russia, or Syria, or Turkmenistan, or Turkey, or Uzbekistan, or Tajikistan, or Czech R, or Slovakia, or Slovenia, or Honduras, or Mexico, or Canada, or United States, or Australia, or Morocco, or Kenya, or Tanzania, or Uganda, or Ethiopia, or Madeira, or New Zealand, or Canary Islands, or South Africa, or Peru, or Ireland, or United Kingdom, or Other Atlantic Islands, or Greece, or Spain, or Portugal, or Albania, or Austria, or Belgium, or Bosnia & Herz, or Bulgaria, or Corsica, or Croatia, or Denmark, or Finland, or France, or Germany, or Hungary, or Iceland, or Italy, or Macedonia, or Norway, or Poland, or Romania, or Sicily, or Sweden, or Switzerland, or Yugoslavia.


Thysanocarpus

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals; white, or purple (or violet); oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); ovate or obovate, or orbicular; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed); fruit appendages wings. Ovules/seeds per ovary/fruit one; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Canada, or Mexico, or United States.


Trachystoma

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; obovate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation segmented.

Morocco.


Transberingia

Beringia R. A. Price et al.

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50, or 51 to 100; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Canada, or Bosnia & Herz, or Belarus, or Bulgaria, or United States.


Trichotolimum

Perennial; herbs, or shrubs or subshrubs; terrestrial; thorns and spines absent; hairs (trichomes) at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves absent; divisions and margins pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals subequalling sepals; white; oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; petal claw and/or filament with hairs or papillae; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina.


Tropidocarpum

Agallis Phil.; Twisselmannia Al-Shehbaz.

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins pinnately lobed (lyrate, runcinate, pinnatifid), or 1–3-pinnatisect. Inflorescences racemes that are bracteate throughout or at least in lower half. Petals distinctly longer than sepals; white; obovate; margin and apex entire. Sepals ascending, or erect but not connivent, or spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or oblong, or triangular; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit lacking or reduced to a narrow rim; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20, or 21 to 50, or 51 to 100; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two, or three to six. Fruit erect to spreading. Fruit segmentation not segmented.

Chile, or Mexico, or United States.


Turritis

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or a mixture of simple and branched; multicellular glands absent; not a weed, or a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; spatulate to oblanceolate; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall woody or corky and thick; dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit more than 100; Seeds biseriate (in two rows); wing present, or absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

Estonia, or Latvia, or Lithuania, or Belarus, or Kyrgyzstan, or Moldova, or Ukraine, or Afghanistan, or Armenia, or Azerbaijan, or China, or Cyprus, or Georgia, or India, or Iran, or Iraq, or Israel, or Japan, or Kazakhstan, or Korea, or Lebanon, or Mongolia, or Nepal, or Pakistan, or Russia, or Syria, or Tajikistan, or Turkmenistan, or Turkey, or Uzbekistan, or Czech R, or Slovakia, or Slovenia, or Canada, or Mexico, or United States, or Australia, or Algeria, or Morocco, or Kenya, or Tanzania, or Uganda, or South Africa, or Lesotho, or Ireland, or United Kingdom, or Rwanda, or Greece, or Portugal, or Spain, or Albania, or Austria, or Bulgaria, or Corsica, or Denmark, or Finland, or France, or Germany, or Switzerland, or Netherlands, or Hungary, or Italy, or Yugoslavia, or Norway, or Poland, or Romania, or Sicily, or Sweden, or Macedonia, or Bosnia & Herz, or Croatia.


Vella

Boleum Desv.; Euzomodendron Coss.; Pseudocytisus Kuntze.

Perennial; shrubs or subshrubs; terrestrial; thorns and spines present, or absent; hairs (trichomes) exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; obovate, or obcordate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens united at least along lower third; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad), or silicle (fruit up to 3 times longer than broad); linear, or oblong; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds), or indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed), or lobes not decurrent; fruit appendages none, or a beaklike style or distal segment. Ovules/seeds per ovary/fruit two, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row), or aseriate (not in rows: when fruit has one or a few seeds); wing present, or absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) present, or absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation segmented.

Algeria, or Morocco, or Spain.


Veselskya

Pyaramidium Boiss.

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; pink (or lavender), or purple (or violet); spatulate to oblanceolate; margin and apex entire. Sepals erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); pyramidal; quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall woody or corky and thick; indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages horns. Ovules/seeds per ovary/fruit three to five; Seeds uniseriate (in one row); wing absent; cotyledon accumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan.


Warea

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul, or auriculate, amplexicaul, or sagittate; divisions and margins entire or repand. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender), or purple (or violet); obovate, or orbicular; margin and apex entire. Sepals spreading, or reflexed; free. Stamens six; petal claw and/or filament with hairs or papillae; filaments of median stamens free; stamens exserted well above sepals and petals. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore at least 1.5 mm long, usually much longer; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

United States.


Weberbauera

Alpaminia O.E. Schulz; Pelagatia O.E. Schulz; Stenodraba O.E. Schulz.

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) a mixture of simple and branched, or exclusively malpighiaceous (2-branched, sessile, medifixed); multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are bracteate throughout or at least in lower half, or racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white, or pink (or lavender); obovate, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything, or petal claw and/or filament with hairs or papillae; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear, or oblong; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit three to five, or six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row), or biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina, or Bolivia, or Chile, or Peru.


Werdermannia

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or a mixture of simple and branched; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals, or subequalling sepals; white, or pink (or lavender); oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals ascending, or erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50, or 51 to 100; Seeds biseriate (in two rows); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) present. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina, or Chile.


Winklera

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base petiolate, or sessile but not auriculate or amplexicaul; divisions and margins 1–3-pinnatisect. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow; obovate, or orbicular; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); orbicular, or cordate or obcordate; angustiseptate (flattened at a right angle to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Afghanistan, or Pakistan, or Tajikistan.


Xerodraba

Skottsbergianthus Boelcke; Skotsbergiella Boelcke.

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves absent; divisions and margins entire or repand. Inflorescences solitary flowers borne on pedicels originating from basal rosette. Petals distinctly longer than sepals; white, or pink (or lavender); oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent, or erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); elliptic, or ovate or obovate; latiseptate (flattened parallel to septum); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit two, or three to five, or six to ten; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Argentina, or Chile.


Yinshania

Cochleariella Y.H. Zhang & R. Vogt; Cochleariopsis Y.H. Zhang; Hilliella (O.E. Schulz) Y.H. Zhang.

Annual or biennial, or perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) exclusively simple, or a mixture of simple and branched; multicellular glands absent; not a weed. Stem leaves present; base petiolate; divisions and margins 1–3-pinnatisect, or trifoliolate, or pinnately compound. Inflorescences racemes that are bracteate throughout or at least in lower half, or racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; white; oblong to elliptic, or spatulate to oblanceolate; margin and apex entire. Sepals spreading; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); oblong, or orbicular, or ellipsoid; angustiseptate (flattened at a right angle to septum), or latiseptate (flattened parallel to septum), or terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing entire (not lobed); fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20; Seeds uniseriate (in one row), or biseriate (in two rows), or aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon accumbent, or incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading, or reflexed (pendulous). Fruit segmentation not segmented.

China, or Vietnam, or Taiwan.


Zerdana

Perennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) at least some dendritic (tree-like); multicellular glands absent; not a weed. Stem leaves absent; divisions and margins entire or repand, or pinnately lobed (lyrate, runcinate, pinnatifid). Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow, or pink (or lavender); obovate, or spatulate to oblanceolate; margin and apex entire. Sepals erect but not connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens united at least along lower third; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes not decurrent; fruit appendages none. Ovules/seeds per ovary/fruit six to ten, or eleven to 20, or 21 to 50; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Iran.


Zilla

Perennial; shrubs or subshrubs; terrestrial; thorns and spines present; hairs (trichomes) absent; multicellular glands absent; not a weed. Stem leaves present; base sessile but not auriculate or amplexicaul; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; pink (or lavender), or purple (or violet); obovate, or oblong to elliptic; margin and apex entire. Sepals ascending; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silicle (fruit up to 3 times longer than broad); didymous (spectacle shaped), or pyriform or obpyriform (pear-shaped); terete (not flattened, cylindrical (circular in cross section)), or quadrangular (4-angled (square or rectangular in cross section)); fruit (or segments) wall woody or corky and thick; indehiscent (seeds remain inside whole fruit); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages wings, or a beaklike style or distal segment. Ovules/seeds per ovary/fruit two; Seeds aseriate (not in rows: when fruit has one or a few seeds); wing absent; cotyledon conduplicate; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Egypt, or Iraq, or Israel, or Jordan, or Kuwait, or Oman, or Qatar, or Saudi Arabia, or Syria, or United Arab Emir, or Yemen, or Algeria, or Libya, or Morocco, or Tunisia, or Mali, or Mauritania.


Zuvanda

Annual or biennial; herbs; terrestrial; thorns and spines absent; hairs (trichomes) absent, or exclusively simple; multicellular glands absent; not a weed. Stem leaves present; base auriculate, amplexicaul, or sagittate; divisions and margins entire or repand, or dentate, serrate, incised, crenate, sinuate. Inflorescences racemes that are ebracteate or rarely lowermost 1 or 2 flowers bracteate. Petals distinctly longer than sepals; yellow, or pink (or lavender), or purple (or violet); obovate; margin and apex entire. Sepals erect and connivent; free. Stamens six; lower part of filaments and petal claws without anything; filaments of median stamens free; stamens included within calyx or partially exserted. Fruit a silique (fruit more than 3 times longer than broad); linear; terete (not flattened, cylindrical (circular in cross section)); fruit (or segments) wall neither woody or papery (normal); dehiscent (readily releasing seeds); gynophore absent or rarely to 1 mm long; septum in mature fruit complete or rarely with a hole; stigma lobing lobes decurrent (connivent); fruit appendages none. Ovules/seeds per ovary/fruit 21 to 50, or 51 to 100; Seeds uniseriate (in one row); wing absent; cotyledon incumbent; seed mucilage (when seeds soaked in water for a few minutes) absent. Leaves alternate. Flowers actinomorphic (all petals equal); All flowers bisexual (with stamens and pistils). Fruit valves two. Fruit erect to spreading. Fruit segmentation not segmented.

Armenia, or Azerbaijan, or Iran, or Iraq, or Israel, or Lebanon, or Saudi Arabia, or Syria, or Turkey, or Egypt.


Cite this publication as: 'Al-Shehbaz, Ihsan (2003). Brassicaceae of the World: Descriptions, Illustrations, Identification, and Information Retrieval. Version: 2003. http://flora.huh.harvard.edu/Brassicaceae/'. Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993, 1995, 2000) should also be cited (see References).

Index